Quedius (Distichalius) iaculifer, Smetana, 2015

Smetana, Aleš, 2015, Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 51. Genus Quedius STEPHENS, 1829. Subgenus Distichalius CASEY, 1915. Section 4, Linzer biologische Beiträge 47 (1), pp. 905-924 : 913-914

Quedius (Distichalius) iaculifer nov.sp. ( Figs 30-36 )

T y p e l o c a l i t y: China:S.GansuWQinlingShan47kmNChengxian, 34° 10’20"N 105

42’19"E, 1830 m.

T y p e m a t e r i a l Holotype (3): CHINA: " CHINA (S Gansu) W Qinling Shan, 47 km N Chengxian 1830 m 34°10’20"N / 105°42’19"E (creek valley with secondary deciduous forest, moist litter sifted) 29.VII.2012 D. V. Wrase [06A]" . Allotype (♀): " CHINA [6]- S-Gansu, N Chengdian , W-Qinling Shan, 34°10’20"N, 105°42’10"E, 1830 m, 29.VII.2012, V. Assing " . Holotype in MSC, allotype in ASC. Paratypes: Gansu: same data as holotype, 533, 5♀♀ ( ASC, MSC): same as holotype, but 34°10’17"N 105° 42’56"E, 1850 m, 29.VII.2012, [ CH 12-05], M. Schülke, 3♀♀ ( ASC, MSC) ; W-Qinling Shan , 43 km N Chengxian, 34°08’16N 105°46’42"E, 1760 m, 28.VII.2012, [CH-12-04], M. Schülke, 1♀ ( MSC) ; same, but 34°10’20"N 105°42’19"E, 1830 m, 29.VII.2012, [ CH 12-06], M. Schülke, 13, 2♀♀ ( MSC) ; same data as allotype, 13, 2♀♀ ( ASC, VAC) ; same, but 34°10’17"N 105°42’56"E, 1850 m, 29.VII.2012, [5], V. Assing, 233, 1♀ ( ASC, VAC) ; same, but 34°08’16"N 105°46’42"E, 1760 m, 28.VII.2012, [4], V. Assing, 233, 3♀♀ ( ASC, VAC). Shaanxi : Qinling Shan, 42 km SW Meixian, 34°01’32"N 107° 24’13"E, 1875 m, 26.VII.2012, [02A], D.W. Wrase, 13 ( MSC) ; same, but [2], V. Assing, 13 ( ASC) .

D e s c r i p t i o n: In all characters similar to Q. quinctius SMETANA, 1998 , but different by a few external characters and particularly by the shape of the aedoeagus and tergite 10 of female genital segment. Average size smaller and body narrower; abdominal tergites piceous-black to black, with apical margins only indistinctly paler; antenna piceous-black with first three segments paler, more or less testaceous, in general shorter, particularly middle segments 4 to 6.

Male. First four segments of front tarsus similar to those of Q. quinctius , but relatively more distinctly dilated. Sternite 8 with two long setae on each side, with medioapical emargination similar to that of Q. quinctius , but somewhat wider and deeper ( Fig. 30 ). Genital segment with tergite 10 narrower, with numerous long setae at and near apex, otherwise asetose ( Fig. 31 ); sternite 9 not appreciably different from that of Q. quinctius . Aedoeagus ( Figs 32-35 ) similar to that of Q. quinctius , including median lobe with apical lobes ( Fig. 33 ), and different mainly by the shape of the paramere, which exceeds with its quite acute apex more distinctly apex of median lobe; number of sensory peg setae on underside of paramere varies to some extent ( Figs 34, 35 ).

Female. First four segments of front tarsus similar to those of Q. quinctius , but somewhat less dilated, segment 2 narrower than apex of tibia (ratio 0.85). Genital segment with tergite 10 wider than that of Q. quinctius , with minute, acute apex, with numerous apical setae of variable length, missing the longitudinal row of fine setae in front of apical setae, present in Q. quinctius ( Fig. 35 and Fig. 28 in SMETANA 1998: 329).

Length 5.8-6.5 mm.

E t y m o l o g y: The specific epithet is a combination of Latin noun iaculum, - i, m (dart, javelin) and the verb fero (to bear, to carry). It refers to the quite acute apex of the paramere of the aedoeagus.

G e o g r a p h i c a l d i s t r i b u t i o n Quedius iaculifer is at present known from several localities in Qinling Shan, both in Gansu and Shaanxi.

B i o n o m i c s Specimenswerecollectedatrelatively lowelevationsbelow 2000 m in a creek valley with secondary deciduous forest by sifting moist forest floor litter, at a margin of a mixed secondary forest by sifting floor litter, and in a secondary deciduous forest on north slope by sifting litter and grass near a creek.

R e c o g n i t i o n a n d c o m m e n t s Quedius iaculifer is also similar to Q. quiris SMETANA, 1998 , but the latter differs, in addition to the different aedoeagus ( Figs. 40-43 in SMETANA 1998: 329), by the more dilated first four segments of the male front tarsus, by the dark maxillary and labial palpi and by the darker three basal segments of the antenna.

Numerous specimens of the original series are teneral, two in bad shape were not included in the paratype series.

Figures

Figs 20-30: 20-23: Quedius causarius sp. nov.: (20) apical portion of aedoeagus, lateral view; (21) apical portion of median lobe, paramere removed; (22) apical portion of underside of paramere with sensory peg setae; (23) tergite 10 of female genital segment. 24-29: Quedius laetepictus sp. nov.: (24) apical portion of male sternite 8; (25) tergite 10 of male genital segment; (26) sternite 9 of male genital segment; (27) aedoeagus, ventral view; (28) apical portion of median lobe, paramere removed; (29) apical portion of underside of paramere with sensory peg setae. Quedius iaculifer sp. nov.: (30) apical portion of male sternite 8.

Figs 31-44: 31-36: Quedius iaculifer sp. nov.: (31) tergite 10 of male genital segment; (32) aedoeagus, ventral view; (33) apical portion of median lobe, paramere removed; (34, 35) apical portion of underside of paramere with sensory peg setae; (36) tergite 10 of female genital segment. 37-42: Quedius wolong: (37) apical portion of male sternite 8; (38) tergite 10 of male genital segment; (39) sternite 9 of male genital segment; (40) aedoeagus, ventral view; (41) apical portion of median lobe, aedoeagus removed; (42) apical portion of underside of paramere with sensory peg setae. Quedius regularis: (43) tergite 10 of female genital segment. Quedius stouraci: (44) tergite 10 of female genital segment.