Anasillomos chrysopos Londt, 1983

Dikow, Torsten, 2015, Review of Anasillomos Londt, 1983 with the description of a new species (Insecta: Diptera: Asilidae), Biodiversity Data Journal 3, pp. 4652-4652 : 4652

Anasillomos chrysopos Londt, 1983

Anasillomos chrysopos Londt, 1983: 284.

Materials

Type status: Holotype. Occurrence: catalogNumber: NMSA-DIP-06957 ; recordedBy: Forchhammer, P.; individualCount: 1; sex: male; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Botswana; stateProvince: Central; locality: Serowe ; verbatimCoordinates: 22°23'00"S 026°43'00"E; decimalLatitude: -22.38333; decimalLongitude: 26.71667; Identification: identifiedBy: Londt, J.; dateIdentified: 1983; Event: eventDate: 1982-08-29; Record Level: institutionCode: NMSA; basisOfRecord: PreservedSpecimen

Type status: Paratype. Occurrence: catalogNumber: NMSA-DIP-66448 ; recordedBy: Forchhammer, P.; individualCount: 1; sex: male; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Botswana; stateProvince: Central; locality: Serowe ; verbatimCoordinates: 22°23'00"S 026°43'00"E; decimalLatitude: -22.38333; decimalLongitude: 26.71667; Identification: identifiedBy: Londt, J.; dateIdentified: 1983; Event: eventDate: 1982-08-29; Record Level: institutionCode: NMSA; basisOfRecord: PreservedSpecimen

Type status: Paratype. Occurrence: catalogNumber: NMSA-DIP-66449 ; recordedBy: Forchhammer, P.; individualCount: 1; sex: male; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Botswana; stateProvince: Central; locality: Serowe ; verbatimCoordinates: 22°23'00"S 026°43'00"E; decimalLatitude: -22.38333; decimalLongitude: 26.71667; Identification: identifiedBy: Londt, J.; dateIdentified: 1983; Event: eventDate: 1982-08-29; Record Level: institutionCode: NMSA; basisOfRecord: PreservedSpecimen

Type status: Paratype. Occurrence: catalogNumber: NMSA-DIP-66450 ; individualCount: 1; sex: male; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Karas; county: Bethanie; locality: Riverside No. 135 ; verbatimCoordinates: 26°38'29"S 016°59'50"E; decimalLatitude: -26.64139; decimalLongitude: 16.99722; Identification: identifiedBy: Londt, J.; dateIdentified: 1983; Event: eventDate: 1971-10-23-1971-10-26; habitat: hover near ground among small shrubs; Record Level: institutionCode: NMSA; basisOfRecord: PreservedSpecimen

Type status: Paratype. Occurrence: catalogNumber: NMSA-DIP-06963 ; individualCount: 1; sex: male; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Karas; county: Bethanie; locality: Riverside No. 135 ; verbatimCoordinates: 26°38'29"S 016°59'50"E; decimalLatitude: -26.64139; decimalLongitude: 16.99722; Identification: identifiedBy: Londt, J.; dateIdentified: 1983; Event: eventDate: 1971-10-23-1971-10-16; habitat: hover near ground among small shrubs; Record Level: institutionCode: NMSA; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: NMSA-DIP-66452 ; recordedBy: Dikow, T.; individualCount: 1; sex: female; lifeStage: adult; previousIdentifications: Anasillomos sp. by Londt, J.; Dikow, T. in 2000; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Karas; locality: Ai-Ais Nature Reserve, NW Gruenau (road D601) ; verbatimCoordinates: 27°30'45"S 017°50'01"E; decimalLatitude: -27.5125; decimalLongitude: 17.83361; Identification: identifiedBy: Dikow, T.; dateIdentified: 2014; Event: eventDate: 1999-12-09; Record Level: institutionCode: NMSA; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: NMSA-DIP-66451 ; recordedBy: Dikow, T.; individualCount: 1; sex: female; lifeStage: adult; previousIdentifications: Anasillomos sp. by Londt, J.; Dikow, T. in 2000; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Karas; locality: Ai-Ais Nature Reserve, NW Gruenau (road D601) ; verbatimCoordinates: 27°30'45"S 017°50'01"E; decimalLatitude: -27.5125; decimalLongitude: 17.83361; Identification: identifiedBy: Dikow, T.; dateIdentified: 2014; Event: eventDate: 1999-12-09; Record Level: institutionCode: NMSA; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: NMSA-DIP-06965 ; recordedBy: Londt, J.; individualCount: 1; sex: female; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: South Africa; stateProvince: Northern Cape; locality: Carnarvon, 2 km NE ; verbatimElevation: 1350 m; verbatimCoordinates: 30°56'07"S 022°07'03"E; decimalLatitude: -30.93528; decimalLongitude: 22.1175; Identification: identifiedBy: Dikow, T.; dateIdentified: 2000; Event: eventDate: 1986-11-14; habitat: flat scrubland; Record Level: institutionCode: NMSA; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: INHS-33959 ; recordedBy: Irwin, M.; individualCount: 1; sex: female; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Karas; county: Karasburg; locality: Ai Ais, 30 km NE, Altdorn No. 3 ; verbatimElevation: 640 m; verbatimCoordinates: 27°48'16"S 017°40'02"E; decimalLatitude: -27.80444; decimalLongitude: 17.66722; Identification: identifiedBy: Dikow, T.; dateIdentified: 2014; Event: samplingProtocol: Malaise trap; eventDate: 1996-11-19; habitat: dry wash; Record Level: institutionCode: INHS; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: individualCount: 1; sex: male; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Karas; locality: Awasib ; verbatimCoordinates: 25°23'00"S 015°39'00"E; decimalLatitude: -25.38333; decimalLongitude: 15.65; Identification: identifiedBy: Londt, J.; dateIdentified: 1983; Event: eventDate: 1971-11-09-1971-11-10; Record Level: institutionCode: NMNW; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: NMNW-H60955 ; recordedBy: Irish, J.; individualCount: 1; sex: female; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Khomas; locality: Windhoek ; verbatimCoordinates: 22°34'00"S 017°05'00"E; decimalLatitude: -22.56667; decimalLongitude: 17.08333; Identification: identifiedBy: Londt, J.; dateIdentified: 2005; Event: eventDate: 1984-09-26; Record Level: institutionCode: NMNW; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: individualCount: 1; sex: female; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Khomas; locality: Windhoek ; verbatimCoordinates: 22°34'00"S 017°05'00"E; decimalLatitude: -22.56667; decimalLongitude: 17.08333; Identification: identifiedBy: Londt, J.; Dikow, T.; dateIdentified: 2000; Event: eventDate: 1973-10-07-1973-10-12; Record Level: institutionCode: NMNW; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: individualCount: 1; sex: undetermined; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Khomas; locality: Windhoek ; verbatimCoordinates: 22°34'00"S 017°05'00"E; decimalLatitude: -22.56667; decimalLongitude: 17.08333; Identification: identifiedBy: Londt, J.; Dikow, T.; dateIdentified: 2000; Event: eventDate: 1973-10-07-1973-10-12; Record Level: institutionCode: NMNW; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: individualCount: 1; sex: female; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Khomas; locality: Windhoek ; verbatimCoordinates: 22°34'00"S 017°05'00"E; decimalLatitude: -22.56667; decimalLongitude: 17.08333; Identification: identifiedBy: Londt, J.; Dikow, T.; dateIdentified: 2000; Event: eventDate: 1973-10-04-1973-10-08; Record Level: institutionCode: NMNW; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: SAM-DIP-A008264 ; recordedBy: SAM Museum Staff; individualCount: 1; sex: male; lifeStage: adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Karas; locality: Great Karas Mountains ; verbatimCoordinates: 27°26'00"S 018°35'22"E; decimalLatitude: -27.43333; decimalLongitude: 18.58944; Identification: identifiedBy: Londt, J.; Event: eventDate: 1936-11-00; Record Level: institutionCode: SAMC; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: USNMENT00995574 ; recordedBy: Moore, A.; individualCount: 1; sex: female; lifeStage: adult; previousIdentifications: Anasillomos sp. by Londt, J. in 1994; Taxon: scientificNameID: urn:lsid:zoobank.org:act:38457B23-1147-4C3D-8263-6E93473A0156; scientificName: Anasillomoschrysopos Londt, 1983; family: Asilidae; genus: Anasillomos; specificEpithet: chrysopos; scientificNameAuthorship: Londt, 1983; Location: country: Namibia; stateProvince: Khomas; locality: Hakos Mountains, 191 km E Walvis Bay ; verbatimCoordinates: 23°14'43"S 016°17'22"E; decimalLatitude: -23.24528; decimalLongitude: 16.28944; Identification: identifiedBy: Dikow, T.; dateIdentified: 2014; Event: eventDate: 1983-11-13; Record Level: institutionCode: USNM; basisOfRecord: PreservedSpecimen

Description

Head: wider than high, black; vertex slightly depressed (less than 60° angle on median margin of compound eye); facial swelling extending over lower ¾ of face, white pubescent; mystax white macrosetose on ventral margin and light brown macrosetose elsewhere, extending over lower ¾ of face; ommatidia of different size, at least some median ommatidia distinctly larger; postgena posterior margin simple, smooth; frons (at level of antennal insertion) more or less parallel-sided, apubescent antero-medially, grey to golden pubescent otherwise, white setose and light brown macrosetose laterally; ocellar tubercle golden pubescent, light brown setose and macrosetose; vertex golden pubescent, asetose; median occipital sclerite (m ocp scl) with several light brown macrosetae; postocular (pocl) setae straight, light brown macrosetae; occiput dorso-medially golden pubescent and laterally grey pubescent, dorso-medially with V-shaped apubescent stripes, white setose.

Proboscis and maxillary palpus: proboscis straight, black; postmentum plate-like, straight, ventral margin entirely smooth, white setose ventrally; prementum circular in cross section proximally, with dorso-median flange, white setose proximo-ventrally; labella reduced, fused to prementum entirely, small, only forming distal tip of proboscis, apex rounded; maxillary palpus two-segmented, black, distal palpomere cylindrical, yellowish setose and macrosetose; stipites fused entirely medially, apubescent, long white setose.

Antenna: light brown, lightly grey pubescent; scape 2x as long as pedicel, short white and light brown setose dorsally and long light brown macrosetose ventrally; pedicel white and light brown setose distally; postpedicel cylindrical (same diameter throughout), 1.5x as long as scape and pedicel combined, sparsely white setose dorso-distally; stylus comprised of 1 element, asetose; apical seta-like sensory element situated apically on stylus.

Thorax: brown to dark brown, postpronotal lobes and lateral scutum light brown to orange; prosternum apubescent, separated from proepisternum, square to rectangular in shape (straight dorsally); proepisternum grey pubescent, white setose and light brown macrosetose; cervical sclerite long white setose; antepronotum golden pubescent, white setose and light brown macrosetose; postpronotum golden pubescent, white setose and light brown macrosetose, postpronotal lobe white setose and long light brown macrosetose; pleuron grey and golden pubescent; proepimeron white setose; anepisternum anterior half asetose, dorso-medially white to light brown macrosetose (6-7 macrosetae), posterior half sparsely long white setose, supero-posteriorly white setose (not macrosetose); anterior basalare asetose, posterior basalare asetose; anepimeron predominantly asetose, antero-dorsally white setose, katepisternum predominantly asetose, postero-dorsally white setose, katepimeron asetose, katatergite white setose and long yellowish macrosetose; meron + metanepisternum asetose, metakatepisternum asetose, metepimeron sparsely white setose; anatergite long white setose; scutum predominantly light brown pubescent, median stripe (extending beyond transverse suture) grey pubescent, paramedian stripes (reaching transverse suture) light brown pubescent, narrow grey pubescent stripes (merging with median stripe beyond transverse suture) grey pubescent, broad stripes (anteriorly wide and posteriorly narrow) brown pubescent (especially visible in anterior and lateral view), sub–lateral spots (1 anterior and 2 posterior to transverse suture) brown pubescent; scutum setation: short white setose anterior and long white setose posterior to transverse suture, setae with small sockets, 3 npl setae, 4 spa setae, 4 pal setae, 2 light brown presutural dc macrosetae and 3-4 light brown postsutural dc macrosetae, acr setae white, presuturally short and longer postsuturally, median posterior scutum (between dc setae) long white setose, setae directed posteriorly; scutellum grey pubescent, ds sctl setae present, long white setae on posterior margin, ap sctl setae present, 6-8 long light brown macrosetae; postmetacoxal area entirely membranous.

Leg: light brown to brown, apubescent, all setae circular in cross section; pro coxa brown, grey pubescent, white setose and light brown macrosetose; pro femur brown, dorsally dark brown, short white setose, light brown macrosetose: 1-2 antero-median, 8-12 postero-dorsal, and 4-5 ventral macrosetae; pro tibia light brown to brown, short white setose, yellowish macrosetose: 7 in 1 antero-dorsal row, 5 in 1 postero-dorsal row, 7 in 1 postero-ventral row, 3 long in 1 postero-ventral row, distally with 6-7 long white macrosetae; mes coxa brown, grey pubescent, white setose and yellowish macrosetose; mes femur brown, dorsally dark brown, short white setose, light brown macrosetose: 2 antero-median, 6-8 postero-dorsal (longest distally), and 5-6 ventral macrosetae; mes tibia brown, short white setose, yellowish macrosetose: 4 in 1 antero-dorsal row, 5 in 1 dorsal row, 5 in 1 posterior row, 3 in 1 postero-ventral row; met coxa brown, grey pubescent, white setose and light brown macrosetose, anteriorly without any protuberance; met trochanter white setose and yellowish macrosetose, cylindrical, medially without protuberance; met femur brown, dorsally dark brown, short white setose, light brown macrosetose: 6-8 anterior, 4-5 ventral, and 3 posterior macrosetae; met tibia brown, straight, short white setose, yellowish macrosetose: 3-4 in 1 antero-dorsal row, 3-4 in 1 dorsal row, 3 in 1 ventral row; proximal pro, mes, and met tarsomeres as long as following 2 tarsomeres combined, pro, mes, and met tarsomeres white setose dorsally, pro tarsomeres white macrosetose disto-laterally and disto-ventrally, mes and met tarsomeres white macrosetose laterally and ventrally; pulvilli well-developed (as long as claw); claw abruptly angled distally, pointed; empodium setiform, well-developed (as long as pulvilli).

Wing: length 8.1-9.5 mm, wing membrane hyaline, without microtrichia; C circumambient (developed around entire wing), anterior wing margin straight; R₂₊₃ distally distinctly arching anteriorly, r₁ open; R₄ terminating anterior to wing apex, distinctly arching anteriorly, stump vein (R₃) absent; r₄ open, R₄ and R₅ diverging from each other; R₅ not reaching C (or wing margin); r₅ closed and petiolate; M₁ not reaching C (or wing margin); cell d closed by base of M₂ and m-m, M₂ and m-m not aligned, r-m situated in distal half of cell d; m₃ closed at C (non-petiolate) or open, M₃ and M₄ approximating at C; cua closed at C (non-petiolate); alula well-developed; microtrichia on posterior wing margin arranged in a single plane.

Abdomen: brown to black, laterally light brown to orange, tergites smooth, setae with small sockets only; T1 white setose, laterally long light brown macrosetose, grey pubescent, entirely sclerotized medially, dorsal surface smooth, without protuberances; T2-8 entirely sclerotized, predominantly black, sometimes brown to orange laterally, predominantly grey pubescent, T2 with apubescent stripe from antero-lateral corner to postero-median margin, T3-5 with narrower apubescent stripes as on T2, white setose, yellowish macrosetae postero-laterally on T2-4 (most pronounced on T2); marginal macrosetae present on T2-4, medial macrosetae present on T2; S1-8 black, lightly grey pubescent, white, erect setose.

Female: T7 and S7 without modifications, ovipositor comprised of 8th and following segments, T6-8 apubescent, setation directed anteriorly on T6-7 and erect on T8; T8 anteriorly with internal rectangular apodeme (entirely fused to T), S8 plate-like, hypogynial valves extending; T9 and T10 entirely fused, sclerites not distinguishable, T10 divided into two heavily sclerotized acanthophorite plates, with 6-7, yellowish acanthophorite spurs per plate; 3 spermathecae, all equally large, reaching anterior end of segment 6; common spermathecal duct short, not extending beyond tip of furca, individual spermathecal ducts long; ejection apparatus absent; spermathecal reservoirs formed by more or less expanded and coiled ducts, weakly sclerotized; furca (S9) formed by single, inverted V-shaped sclerite, median sclerite (at posterior tip) absent, anterior furcal apodeme absent.

Male (Fig. 4a, b, c): T1-T8 and S1-S8 entire (without modifications); hypopygium dark brown, rotated by 180°, directed posteriorly; epandrium divided medially into two halves, joined proximally; hypandrium well-developed, triangular, distally with short postero-median projection, distinctly separated from epandrium by gonocoxite, not fused to gonocoxite; gonocoxite partially fused to epandrium proximally, gonocoxal apodeme present, short, at most slightly extending hypopygium proximally; gonostylus present, positioned distally on gonocoxite; subepandrial sclerite asetose, ventrally with median, wavy longitudinal ridge, laterally straight (without protuberances), distally simple, straight margin; cerci separate (not fused medially); 1 aedeagal prong, tip pointed, without any protuberance, dorsal aedeagal sheath short, sperm sac entirely free; lateral ejaculatory process present, large triangular sclerite, free (not surrounded by ventral aedeagal sheath); ejaculatory apodeme formed by single vertical plate (two lateral surfaces).

Diagnosis

The species is distinguished from its congener by the dorso-medially white macrosetose anepisternum (in addition to white setae, Figs 2b, 3b), the distinct pubescence pattern on the lateral abdominal tergites (T2-5 with apubescent stripe from antero-lateral corner to postero-median margin, Fig. 3b), and the presence of long macrosetae on the tip of the pointed hypandrium (almost reaching tip of epandrium, Fig. 4b).

Distribution

Known from Botswana, Namibia, and South Africa and therefore widely distributed in southern Africa, but rarely collected (Fig. 11).

Taxon discussion

The type locality of A. chrysopos lies in eastern Botswana, far removed from the majority of specimens collected in western Namibia. There is little morphological variation within the species except for the setation color, which can vary from light brown to yellow.

Type locality

Botswana: Central: Serowe (22°23'00"S 026°43'00"E).

Biodiversity hotspot

Not known to occur in any of the southern African biodiversity hotspots (Cape Floristic Region, Maputaland-Pondoland-Albany, or Succulent Karoo).