Aulacopalpus pygidialis Ohaus, 1905

Smith, Andrew B. T., 2002, Revision Of The Southern South American Endemic Genus Aulacopalpus Guérin-Méneville With Phylogenetic And Biogeographic Analyses Of The Subtribe Brachysternina (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini), The Coleopterists Bulletin 56 (3), pp. 379-437 : 414-417

Aulacopalpus pygidialis Ohaus, 1905

( Figs. 1 , 7 , 19 , Map 3)

Aulacopalpus pygidialis Ohaus 1905:139 . Lectotype male at the ZMHB, labeled a) ‘‘ CHILE,’’ b) ‘‘Typus!,’’ c) ‘‘ A. pygidialis Ohaus ,’’ d) ‘‘AU­ LACOPALPUS PYGIDIALIS OHAUS Ƌ DET: A. B. T. SMITH 2000 LECTOTYPE,’’ e) ‘‘Zool. Mus. Berlin.’’ LECTOTYPE HERE DES­ IGNATED. See statement of taxonomic purpose in Methods and Materials section. The existence and location of other male paralectotypes is unknown (Ohaus had no female specimens).

Description. Male (n = 22): Length 14.6–18.5 mm. Width 7.6–9.5 mm. Color: dorsally brown to greenish­brown, head and pronotum dark brown to brown with greenish reflections; venter brown. Head: Dorsal surface with margin sparsely setose; setae long, slender, tawny to cream­colored. Frontoclypeal suture weakly bisinuate. Clypeal surface brown, rugose. Frons green to brown with greenish reflections, densely punctate. Labrum moderately produced at middle, rounded. Maxillary palpus with terminal segment slightly shorter than basal 3 segments, with sulcus. Mentum with surface moderately setose; apex reflexed into oral cavity. Antenna 10­segmented, club greatly elongated (longer than combined basal segments), flattened. Pronotum: Midline weakly punctate. Surface moderately to densely punctate and moderately setose; punctures moderately large; setae long, slender, tawny to cream­colored. Scutellum: Shape parabolic: 1.2 times wider than long medially. Surface densely punctate, moderately setose; punctures moderately large; setae long, slender, tawny to cream­colored. Elytron: Surface glabrous; longitudinal striae punctate, weakly impressed; punctures moderately­sized, separated by 1–4 puncture widths; intervals sparsely punctate with moderately­sized punctures. Epipleuron with row of dense setae just ventral to bead present medially, absent in apical and basal 1/5. Pygidium: Width 1.6 times length medially. Surface moderately punctate, moderately setose; punctures moderate­sized; setae tawny. Venter: Thorax densely setose; setae long, tawny. Legs: Protibia with 3 subequally large teeth in apical half. Protarsal claws ( Fig. 7 ) with modified claw slightly thickened when compared with other claw, apex weakly bifurcate (often worn, appearing simple). Meso­ and metatarsal claws simple (similar to Fig. 6 ). Tarsomere 5 with ventromedial tooth ( Fig. 7 ). Meso­ and metatibia with apical spurs slender, acute. Mesotibial apex with 12–13 weak spinules. Metatibial apex with 18–20 weak spinules. Parameres: Figure 19 . Basally without broad, medial furrow.

Female (n = 1): Length 16.4 mm. Width 8.2 mm. As male except in the following respects. Head: Antennal club not elongated and flattened, equal to or less than length of terminal segments. Legs: Tarsal claws simple. Meso­ and metatibia with apical spurs broader, blunter.

Diagnosis. This species is distinguished from all other species in the genus Aulacopalpus by the following combination of characters: antennal club elongated (length equal or slightly shorter than head); head and pronotum dark brown to brown with greenish reflections; elytra brown to greenish brown; male protarsal claws with modified claw slightly thickened when compared with other claw and with apex weakly bifurcate (often worn and appearing simple); female protarsal claws simple; meso­ and metatarsal claws simple in both sexes; and tarsomere 5 with ventromedial tooth; parameres basally without broad, medial furrow.

Distribution (Map 3). Chile from Concepción to Llanquihue.

Locality Data. Based on 23 specimens examined from CMNC, FMNH, MNNC, ZMHB .

THE COLEOPTERISTS BULLETIN 56(3), 2002 415

Map 3. Distribution of Aulacopalpus pygidialis (O), Aulacopalpus valdiviensis (*), and Aulacopalpus viridis (•).

416 THE COLEOPTERISTS BULLETIN 56(3), 2002

Map 4. Range of Brachysternina showing north and south limits to Nothofagus (after

Veblen et al. 1996), Aulacopalpus , Brachysternus , and Hylamorpha .

THE COLEOPTERISTS BULLETIN 56(3), 2002 417

CHILE (21). CONCEPCIÓN (1): Concepción. LLANQUIHUE (8): Llanquihue, Maullín. OSORNO (11): Pucatrihue Costa. NO DATA (1).

NO DATA (2).

Temporal Data. August (11), September (3), October (1), November (5).

Remarks. Ohaus (1905) described Aulacopalpus pygidialis using only male specimens. This is the first time the female has been described.

Figures

Fig. 1. The Brachysternina. First row: Aulacopalpus aconcaguensis (male paratype), A. aconcaguensis (male paratype), A. castaneus (male), A. castaneus (male), A. castaneus (male), A. punctatus (male). Second row: A. ciliatus (male), A. clypealis (male),

Figs. 6–11. 6) Protarsomere 5 and claw of male Aulacopalpus pilicollis, 7) Protarsomere 5 and claw of male Aulacopalpus pygidialis. 8) Protarsomere 5 and claw of female Aulacopalpus ciliatus. 9) Dorsal view of modified proclaw of male Aulacopalpus ciliatus. 10) Dorsal view of modified proclaw of male Aulacopalpus castaneus. 11) Lateral view of Aulacopalpus valdiviensis parameres and phallobase.

Figs. 12–20. Parameres in caudal view. 12) Aulacopalpus aconcaguensis; 13) Aulacopalpus ciliatus; 14) Aulacopalpus clypealis; 15) Aulacopalpus pilicollis; 16) Aulacopalpus castaneus; 17) Aulacopalpus punctatus; 18) Aulacopalpus viridis; 19) Aulacopalpus pygidialis, 20) Aulacopalpus valdiviensis.