Drymaria Willd.

Valentín-Martínez, Dagoberto, Steinmann, Victor W., Flores-Olvera, Hilda, Borsch, Thomas & Montesinos-Tubée, Daniel B., 2025, A nomenclator of Drymaria (Caryophyllaceae), PhytoKeys 266, pp. 139-175 : 139-175

1. Drymaria Willd. in Roem. & Schult., Syst. Veg. 5: 406. 1819. Type (designated by Fosberg 1945, pg. 96): Drymaria cordata (L.) Willd.

Pinosia Urb., Arkiv för Botanik 23 A (5): 70. 1930. Type: Pinosa ortegioides (Griseb.) Urb. (= Drymaria ortegioides Griseb. )

Mollugophytum M. E. Jones, Contr. W. Bot. 18: 35 (1933). Type, designated here: Mollugophytum holosteoides (Benth.) M. E. Jones (= Drymaria holosteoides Benth. )

Notes.

Although authorship of Drymaria is often given as “ Willd. ex Schult., ” the name and description occur in a single paragraph followed by “ Reliq. Willd. MS. ” This represents direct association of Willdenow with both the name and the description, and Willdenow should be cited as author (see Turland et al. (2018), art. 46.3 example 15). Drymaria arenarioides has been stated to be the type of Drymaria (e. g., Duke 1961, Hernández-Ledesma et al. 2015), with Britton and Millspaugh (1920) credited with having made the designation. However, according to Richard Rabeler (pers. comm. 2025), Britton and Millspaugh followed the largely mechanical method of choosing the first species as the type, and their choice is superseded by Fosberg’s designation of Drymaria cordata as the type (see Turland et al. (2018) art. 10.7), which was made in the context of discussing morphological characters to evaluate the status of Mollugophytum and did not follow the mechanical method.

Mollugophytum holosteoides is here designated the type because it is the most widespread species of those treated by Jones (1933) within Mollugophytum .