Guithone, Zheng & Ren & Wang, 2016

Zheng, Bingyu, Ren, Dong & Wang, Yongjie, 2016, Earliest true moth lacewing from the Middle Jurassic of Inner Mongolia, China, Acta Palaeontologica Polonica 61 (4), pp. 847-851 : 849

publication ID

https://doi.org/ 10.4202/app.00259.2016

persistent identifier

https://treatment.plazi.org/id/641E8787-FFDF-FFCD-B0C9-F4D8FCB8F94D

treatment provided by

Felipe

scientific name

Guithone
status

 

Genus Guithone View in CoL nov.

Etymology: From Chinese gu, ancient; and the extant genus name Ithone .

Type species: Guithone bethouxi gen. et sp. nov., monotypic, see below.

Diagnosis. —The same as for the monotypic type species.

Remarks. —Although the specimen of the type species of this genus is only sub-complete, useful characters could be investigated. The new genus is assigned to the Ithonidae according to the following characters: stout body, head hypognathous and concealed under pronotum, and extremely broad costal space at the base ( Fig. 1A 1 View Fig , B). To determine the systematic position of Guithone gen. nov. within Ithonidae , a comparison was conducted between the new genus and the three main lineages ( Table 1). It is noticeable that a simple humeral veinlet (irrecurrent) occurs in the new genus, in contrast to the other genera of Ithonidae that commonly possess the recurrent humeral veinlet. Makarkin et al. (2013) pointed that the recurrent humeral veinlet as an apomorphy of Neuroptera independently evolves in different families within this order. As for Ithonidae , the recurred humeral veinlet should be a possible synapomorphy of this family considering the general occurrence at most known genera. Essentially, the humeral veinlet of Guithone gen. nov. also exhibits a “recurrent trend” in comparison with other genera ( Figs. 2A, B View Fig , 3A). Consequently we consider that the form of humeral veinlet in Guithone gen. nov. represents an intermediate state in the evolution of the character within the Ithonidae .

The Table 1 summarized the distribution of wing venation character states among three genus-groups of Ithonidae , as compared to the states exhibited by Guithone gen. nov. It shows that the new genus shares most character states with the ithonid genus-group. Therefore, we propose that Guithone gen. nov. should be assigned to this group, hence representing its eldest record of this lineage, as well as the first record in the Northern Hemisphere.

Stratigraphic and geographic range.—As for the type species by monotypy.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Neuroptera

Family

Ithonidae

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