Glyphidrilus malayanus Michaelsen, 1902

Chanabun, Ratmanee, Sutcharit, Chirasak, Tongkerd, Piyoros & Panha, Somsak, 2013, The semi-aquatic freshwater earthworms of the genus Glyphidrilus Horst, 1889 from Thailand (Oligochaeta, Almidae) with re-descriptions of several species, ZooKeys 265, pp. 1-76 : 12-14

6. Glyphidrilus malayanus Michaelsen, 1902 Fig. 8 Table 1

Glyphidrilus malayanus Michaelsen, 1902: 35.Type locality: Malay Peninsula, Lubock Paku, Pahang River. Jamieson 1968: 393. Brinkhurst and Jamieson 1971: 762, fig. 15.4E, F. Shen and Yeo 2005: 19. Chanabun et al. 2011: 215, 2012a: 268.

Material examined.

The specimen with the measurements and anatomical characters agreeing with the original description is designated as the lectotype ZMH V5875 ( Fig. 8 ). The type locality of this species is Malay Peninsula, Lubock Paku, Pahang River. Paralectotype: ZMH V5875.1 (only one tail) same locality with lectotype.

Lectotype designation proposed herewith is necessary to ensure the species is based on a complete adult earthworm.

Description of Lectotype.

Dimensions: body length 87 mm. Body cylindrical in anterior part, quadrangular in transverse section-behind clitellum. 246 segments. Body colour pale brown. At posterior end dorsal surface considerably broader than the ventral. Clitellar wi on ventro-lateral part of clitellum in ¾18-½22. Prostomium zygolo bous. Dorsal pores absent. Clitellum annular in 15-26. Four pairs of setae per segment from 2, setal formula aa:ab:bc:cd:dd = 1.0:0.5:1.5:0.5:1.0 in segment 8. Female pores, male pores and spermathecal pores not visible. Gm laterally paired or asymmetrical on bc in 15-17.

Septa 5/6-7/8 thicker, 8/9-11/12 thick and 12/13 to the last segment thin. Gi small, globular within 8. Intestine enlarged from 14. Dorsal blood vessel anterior to 9. He in 9-11. Np distinguishable from segment 17 onwards. Sv in 9-12, those of segment 12 are larger than the others. Ov in 13-14. Testis in 10-11. Prostate and accessory glands absent. Sc in 14/15-16/17, two on each side per segment.

Remarks.

See Table 1 .

The distribution of Glyphidrilus malayanus is Malay Peninsula, Lubock Paku, Pahang River. Sungei Kelatan, Kelantan, Malaysia. Sungei Berkoh, Johor, Malaysia. Tok Bok Hotspring, Kelantan, Malaysia.

Figures

Figure 8. Morphology of the lectotype (ZMH V 5875) of Glyphidrilus malayanus Michaelsen, 1902, showing the A external ventral and B internal dorsal views.