Seeversiella Ashe, 1986

Gusarov, Vladimir I., 2003, A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae), Zootaxa 142, pp. 1-102 : 3-9

Seeversiella Ashe, 1986 ( Figs. 1­384 )

A new genus: Seevers, 1978: 46 (unnamed new genus appearing in a key). Seeversiella Ashe : 1986: 501 (tribe Athetini Casey, 1910 ).

Seeversiella: Ashe in Newton, Thayer, Ashe & Chandler, 2000: (tribe Athetini , not assigned to subtribe).

Diagnosis. Seeversiella can be distinguished from other athetine genera by the combination of the following characters: body parallel­sided or with broad ovate abdomen ( Figs. 17­19 ); antennal articles 8­10 transverse ( Fig. 9 ); ligula divided into two separate lobes ( Fig. 6 ); pronotum with microsetae directed posteriorly along the midline; in lateral portions of the disc microsetae directed towards the midline and/or obliquely posteriorly ( Figs. 10­12 ) (posteriorly in S. geostiboides ); pronotal macrosetae short; pronotal hypomera fully visible in lateral view; medial macroseta of mesotibia inconspicuous, shorter than tibial width; tarsal formula 4­5­5; metatarsal segment 1 as long as segment 2; one empodial seta; in many species posterior angles of male tergum 3 projecting as spines ( Figs. 15 , 17 ), long in large males, short or absent in small males ( Figs. 10 A­10C in Ashe (1986)); in many species male tergum 7 with medial carina along midline ( Figs. 17­18 ) (absent in small males); copulatory piece of internal sac of aedeagus long and flagellumlike ( Figs. 29, 36 ; CP).

Seeversiella differs from Geostiba in having microsetae of lateral portions of pronotum directed towards the midline; posterior angles of male tergum 3 projecting as spines; and long flagellum­like copulatory piece of internal sac of aedeagus.

Seeversiella differs from Tropimenelytron Pace, 1983 in having contiguous mesocoxae; ligula divided into two separate lobes; microsetae of lateral portions of pronotum directed towards the midline; posterior angles of male tergum 3 projecting as spines; and long flagellum­like copulatory piece of internal sac of aedeagus.

Seeversiella differs from the small species of Atheta Thomson, 1858 with transverse antennal segments in having ligula divided into two separate lobes; microsetae of lateral portions of pronotum directed towards the midline; posterior angles of male tergum 3 projecting as spines; and long flagellum­like copulatory piece of internal sac of aedeagus.

Description. Length 1.6­3.7 mm, pronotal width 0.34­0.74 mm. Body parallel­sided or with broad ovate abdomen ( Figs. 17­19 ), dark brown to brownish yellow.

Head as wide as long; eyes small or large, temple length to eye length ratio 0.8­7.0; infraorbital carina incomplete. Antennal article 2 longer or as long as article 3, articles 4­5 slightly elongate, subquadrate or slightly transverse, 6­7 subquadrate or transverse, 8­10 transverse or strongly transverse (ratio 1.5­2.0), apical article without coeloconic sensilla. Labrum ( Fig. 1 ) transverse, anterior margin slightly concave. Adoral surface of labrum (epipharynx) as in Fig. 2 . Mandibles ( Figs. 4­5 ) broad, right mandible with a small medial tooth; dorsal molar area with velvety patch consisting of very small denticles (not visible at 400x). Maxilla ( Fig. 3 ) with galea extending beyond apex of lacinia; apical lobe of galea covered with numerous fine and short setae; apical fifth of lacinia with row of closely spaced spines, middle portion produced medially and covered with numerous setae. Labium as in Figs. 6­8 ; ligula divided into two lobes; medial area of prementum without pores but with 5­15 pseudopores, lateral areas with 3 (occasionally 4) pores and single spinose pore ( Fig. 6 ). Hypopharyngeal lobes as in Fig. 7 . Mentum ( Fig. 8 ) with slightly protruding anterior angles and straight anterior margin.

Pronotum ( Figs. 10­12 ) slightly transverse, broadest near middle, sides slightly convex; anterior and posterior margins convex; surface covered with microsetae directed posteriorly in midline; in lateral areas of the disc microsetae directed towards midline and/or obliquely posteriorly (posteriorly in S. geostiboides ); macrosetae short; hypomera fully visible in lateral view. Meso­ and metasternum as in Fig. 12 , mesosternal process extending about ½ length of mesocoxal cavities, metasternal process short, mesosternum and mesosternal process not carinate medially; relative lengths of mesosternal process: isthmus: metasternal process in ratio of about 3:3:1; mesocoxal cavities margined posteriorly; mesocoxae contiguous. Medial microseta of mesotibia inconspicuous, shorter than tibial width. Tarsal segmentation 4­5­5, metatarsal segment 1 as long as segment 2. One empodial seta, slightly shorter than claws. Wings fully developed, partially reduced or reduced to short vestiges that are shorter than elytra. Posterior margin of elytra straight or slightly concave near postero­lateral angle.

Abdominal terga 3­5 with moderate basal impressions. Tergum 7 as long as tergum 6. Punctation on terga 6­7 sparser than on terga 3­5. Tergum 7 with wide white palisade fringe or without fringe (in wingless species).

In many species posterior angles of male tergum 3 projecting as spines ( Figs. 15 , 17 ), long in large males, short or absent in small males ( Figs. 10 A­10C in Ashe (1986)); in some species lateral portions of posterior margin of male tergum 3 extending as short and obtuse projections ( Figs. 16 , 18­19 ); in most species male tergum 7 with medial carina along midline ( Figs. 17­18 ) (absent in small males) or with medial subapical tubercle. Copulatory piece of internal sac of aedeagus long and flagellum­like ( Figs. 29, 36 ; CP); medial lamellae narrow ( Figs. 29­30 ; 47; 258; ML). Internal sac with one pair of distal sclerites (not homologous to suspensoria) laterally of copulatory piece; the distal sclerites are hook­shaped ( Figs. 28­29, 31, 35 ), dentiform ( Figs. 63, 65 ) or reduced to elongate ( Figs. 199­200, 203, 206 ) or subquadrate ( Fig. 343 ) plates which may have numerous spicules ( Figs. 99­101 ). Spermatheca short, S­, L­, J­ or C­shaped ( Figs. 32 , 53 , 62 , 104 , 149 , 205 ).

Type species. Seeversiella bispinosa Ashe, 1986 , by original designation.

Discussion. Although most species of Seeversiella described below can be recognized by comparing the external shape of the aedeagus, the details of the internal sac provide additional characters to distinguish the closely related species. Some characters of the internal sac, like long flagellum­like copulatory piece ( Figs. 29 ; 172; 200, 203­204; 301; CP) and narrow medial lamellae ( Figs. 29­30 ; 274­275; 304­305; ML), are consistent within the genus. The base of the copulatory piece and the medial lamellae are linked together by two connecting bands ( Figs. 243­245 ; CB). These bands are sclerotized areas of the internal sac wall. In everted internal sac the bands are situated on the parameral face of the sac ( Fig. 245 ; CB). When the sac is retracted the bands are clearly visible laterally of the copulatory piece ( Fig. 243 ; CB).

Other elements of the internal sac may differ significantly between species. One such element is a pair of distal (in everted sac) sclerites located laterally of the copulatory piece ( Figs. 28­29 ; DS). These sclerites are not attached to the copulatory piece and probably they are not homologous to suspensoria. In S. globicollis and S. texana the distal sclerites are hook­shaped ( Figs. 31 ; 51) and, when retracted, lie in the separate pockets of the internal sac. In many species the distal sclerites are reduced to elongate ( Figs. 199­200, 203, 206 ) or subquadrate ( Fig. 343 ) plates which may have numerous spicules ( Figs. 99­101 ). The homology of these plates to the hook­shaped sclerites of S. globicollis can be established by their similar position in the internal sac, laterally of the copulatory piece ( Figs. 28­29 ; 199­200, 203­204). Another detail of the internal sac which varies within the genus is the lateral diverticula which may be covered with numerous setae ( Figs. 29 ; 169, 172­ 173; LD), have sclerotized denticles ( Fig. 200 ) or possess long spiniform sclerites which may look like the medial lamellae but their location in the internal sac is different ( Figs. 301, 304 ; 318; SLD). In retracted sac, the setae, the denticles or the sclerites of the lateral diverticula are usually well visible in cleared preparations ( Figs. 173 ; 299, 303; 316; 327).

In Seeversiella , as in many other aleocharine genera (e.g., Geostiba , Tropimenelytron ) the male secondary sexual characters are subject to intraspecific variation. Larger males have longer spines on male tergum 3 (up to 3 times as long as tergum medial length) and longer carina on tergum 7. In smaller males the spines are very short or absent altogether. Therefore the male secondary characters cannot be reliably used to distinguish the species of Seeversiella .

My examination of available specimens of Seeversiella suggests that some species differ from each other in the length of elytra and wings, and in the eye size. On several occasions these characters are used below in the key to species. However, these characters should be used with caution because some species are polymorphic. Ultimately the details of male genitalia are the most reliable characters to recognize the species of Seeversiella .

Figures

FIGURES 1 ­ 5. Mouthparts of Seeversiella globicollis (Bernhauer) (Waterton Lakes National Park, Alberta). 1 – labrum; 2 – epipharynx; 3 – right maxilla, ventral view; 4 – left mandible, dorsal view; 5 – right mandible, dorsal view. Scale bar 0.1 mm.

FIGURES 6 ­ 9. Mouthparts and antenna of Seeversiella globicollis (Bernhauer) (6 ­ 7, 9, Waterton Lakes National Park, Alberta; 8, Anticosti, Quebec). 6 – prementum; 7 – hypopharynx; 8 – mentum; 9 – right antenna. Scale bar 0.1 mm (6 ­ 8), 0.2 mm (9).

FIGURES 10 ­ 16. Details of Seeversiella globicollis (Bernhauer) (10, Anticosti, Quebec; 13, Waterton Lakes National Park, Alberta), S. scabricollis Gusarov, sp. n. (11, 15, paratype from Volcán Barva, Costa Rica) and S. micralymma Gusarov, sp. n. (12, 14, 16, paratype from Cerro Buenavista, Costa Rica). 10 ­ 12 – pronotum; 13 ­ 14 – mesometathorax; 15 – male tergum 3 and paratergites; 16 – male tergum 3. Scale bar 0.4 mm (10 ­ 11, 15), 0.3 mm (12 ­ 13, 16), 0.2 mm (14).

FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm.

FIGURES 20 ­ 23. Abdominal segment 8 of Seeversiella globicollis (Bernhauer) (20 ­ 21, Cloudcroft, New Mexico; 22 ­ 23, Salmo, British Columbia). 20 – male tergum 8; 21 – male sternum 8; 22 – female tergum 8; 23 – female sternum 8. Scale bar 0.4 mm.

FIGURES 24 ­ 27. Aedeagus of Seeversiella globicollis (Bernhauer) (Cypress Hills Provincial Park, Alberta). 24 – median lobe, parameral view; 25 – apex of median lobe, parameral view; 26 – median lobe, lateral view; 27 – apex of median lobe, lateral view. Scale bar 0.2 mm (24, 26), 0.1 mm (25, 27).

FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 – details of internal sac retracted into median lobe, abparameral view; 29 – everted internal sac, parameral view; 30 – medial lamellae, parameral view; 31 – right hook of internal sac; 32 – spermatheca; 33 – apex of left paramere; 34 ­ 35 – details of internal sac retracted into median lobe, lateral view; 36 – copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36).

FIGURES 37 ­ 40. Abdominal segment 8 of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 37 – male tergum 8; 38 – male sternum 8; 39 – female tergum 8; 40 – female sternum 8. Scale bar 0.4 mm.

FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 – median lobe, parameral view; 42 – apex of median lobe, parameral view; 43 – median lobe, lateral view; 44 – apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44).

FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52).

FIGURES 54 ­ 57. Abdominal segment 8 of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 54 – male tergum 8; 55 – male sternum 8; 56 – female tergum 8; 57 – female sternum 8. Scale bar 0.2 mm.

FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66).

FIGURES 68 ­ 71. Abdominal segment 8 of Seeversiella liliputana Gusarov, sp. n. (68 ­ 69, holotype; 70 ­ 71, possible female of S. liliputana from Bear Saddle, Arizona). 68 – male tergum 8; 69 – male sternum 8; 70 – female tergum 8; 71 – female sternum 8. Scale bar 0.2 mm.

FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 – median lobe, parameral view; 73 – apex of median lobe, parameral view; 74 – median lobe, lateral view; 75 – apex of median lobe, lateral view; 76 – details of internal sac retracted into median lobe, lateral view; 77 – apex of left paramere; 78 – spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78).

FIGURES 79 ­ 82. Abdominal segment 8 of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 79 – male tergum 8; 80 – male sternum 8; 81 – female tergum 8; 82 – female sternum 8. Scale bar 0.2 mm.

FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89).

FIGURES 90 ­ 93. Abdominal segment 8 of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (90 ­ 91) and Las Piedrecitas (92 ­ 93), Mexico). 90 – male tergum 8; 91 – male sternum 8; 92 – female tergum 8; 93 – female sternum 8. Scale bar 0.2 mm.

FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104).

FIGURES 105 ­ 108. Abdominal segment 8 of Seeversiella tuberculicauda Gusarov, sp. n. (105 ­ 106, holotype; 107 ­ 108, paratype from Jalapa, Mexico). 105 – male tergum 8; 106 – male sternum 8; 107 – female tergum 8; 108 – female sternum 8. Scale bar 0.2 mm.

FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115).

FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 – male tergum 8; 118 – male sternum 8; 119 – median lobe, parameral view; 120 – apex of median lobe, parameral view; 121 – median lobe, lateral view; 122 – apex of median lobe, lateral view; 123 ­ 124 – details of internal sac retracted into median lobe, lateral view; 125 – apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS – distal sclerite of internal sac.

FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 – male tergum 8; 127 – male sternum 8; 128 – median lobe, parameral view; 129 – apex of median lobe, parameral view; 130 – median lobe, lateral view; 131 – apex of median lobe, lateral view; 132 – details of internal sac retracted into median lobe, abparameral view; 133 – details of internal sac retracted into median lobe, lateral view; 134 – apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134).

FIGURES 135 ­ 138. Abdominal segment 8 of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (135 ­ 136) and Suchixtepec (137 ­ 138), Mexico). 135 – male tergum 8; 136 – male sternum 8; 137 – female tergum 8; 138 – female sternum 8. Scale bar 0.2 mm.

FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142).

FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149).

FIGURES 150 ­ 153. Abdominal segment 8 of Seeversiella badia Gusarov, sp. n. (150 ­ 151, holotype; 152 ­ 153, paratype, 21.7 km NE Metapan, El Salvador). 150 – male tergum 8; 151 – male sternum 8; 152 – female tergum 8; 153 – female sternum 8. Scale bar 0.2 mm.

FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160).

FIGURES 161 ­ 164. Abdominal segment 8 of Seeversiella similis Gusarov, sp. n. (161 ­ 162, holotype; 163 ­ 164, paratype from La Tigra, Honduras). 161 – male tergum 8; 162 – male sternum 8; 163 – female tergum 8; 164 – female sternum 8. Scale bar 0.2 mm.

FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 – median lobe, parameral view; 166 – apex of median lobe, parameral view; 167 – median lobe, lateral view; 168 – apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168).

FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174).

FIGURES 175 ­ 178. Abdominal segment 8 of Seeversiella minima Gusarov, sp. n. (175 ­ 176, holotype; 177 ­ 178, paratype, 21.7 km NE Metapan, El Salvador). 175 – male tergum 8; 176 – male sternum 8; 177 – female tergum 8; 178 – female sternum 8. Scale bar 0.2 mm.

FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189).

FIGURES 190 ­ 193. Abdominal segment 8 of Seeversiella scabricollis Gusarov, sp. n. (190 ­ 191, holotype; 192 ­ 193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 – male tergum 8; 191 – male sternum 8; 192 – female tergum 8; 193 – female sternum 8. Scale bar 0.4 mm.

FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 – median lobe, parameral view; 196 – apex of median lobe, parameral view; 197 – median lobe, lateral view; 198 – apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198).

FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207).

FIGURES 208 ­ 211. Abdominal segment 8 of Seeversiella furcativentris Gusarov, sp. n. (208 ­ 209, holotype; 210 ­ 211, paratype, Zurquí de Moravia, Costa Rica). 208 – male tergum 8; 209 – male sternum 8; 210 – female tergum 8; 211 – female sternum 8. Scale bar 0.2 mm.

FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 – median lobe, parameral view; 213 – apex of median lobe, parameral view; 214 – apex of median lobe, lateral view; 215 – median lobe, lateral view; 216 – apex of left paramere, 217 – details of internal sac retracted into median lobe, abparameral view; 218 – details of internal sac retracted into median lobe, lateral view; 219 – right distal sclerite of internal sac; 220 – left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 – spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221).

FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 – male tergum 8; 223 – male sternum 8; 224 – female tergum 8; 225 – female sternum 8. Scale bar 0.2 mm.

FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229).

FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234).

FIGURES 235 ­ 238. Abdominal segment 8 of Seeversiella brunnea Gusarov, sp. n. (235 ­ 236, holotype; 237 ­ 238, paratype, P. N. Tapantí, Costa Rica). 235 – male tergum 8; 236 – male sternum 8; 237 – female tergum 8; 238 – female sternum 8. Scale bar 0.2 mm.

FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 – median lobe, parameral view; 240 – apex of median lobe, parameral view; 241 – median lobe, lateral view; 242 – apex of median lobe, lateral view; 243 – details of internal sac retracted into median lobe, lateral view; 244 – apex of left paramere; 245 – everted internal sac, parameral view; 246 – spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246).

FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 – male tergum 8; 248 – male sternum 8; 249 – female tergum 8; 250 – female sternum 8. Scale bar 0.4 mm.

FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255).

FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 – details of internal sac retracted into median lobe, abparameral view; 257 – details of internal sac retracted into median lobe, parameral view; 258 – medial lamellae, parameral view; 259 – everted internal sac, parameral view; 260 – apex of left paramere; 261 ­ 262 – details of internal sac retracted into median lobe, lateral view; 263 – right distal sclerite of internal sac; 264 – everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263).

FIGURES 265 ­ 268. Abdominal segment 8 of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 265 – male tergum 8; 266 – male sternum 8; 267 – female tergum 8; 268 – female sternum 8. Scale bar 0.2 mm.

FIGURES 269 ­ 277. Genitalia of Seeversiella lativentris Gusarov, sp. n. (paratypes from Villa Mills, Costa Rica). 269 – median lobe, parameral view; 270 – apex of median lobe, parameral view; 271 – median lobe, lateral view; 272 – apex of median lobe, lateral view; 273 – apex of left paramere; 274 – everted internal sac, parameral view; 275 – everted internal sac, lateral view; 276 – details of internal sac retracted into median lobe, lateral view; 277 ­ spermatheca. Scale bar 0.2 mm (269, 271, 274 ­ 276), 0.1 mm (270, 272 ­ 273, 277).

FIGURES 278 ­ 281. Abdominal segment 8 of Seeversiella luridicollis Gusarov, sp. n. (paratypes from Cerro Buenavista, Costa Rica). 278 – male tergum 8; 279 – male sternum 8; 280 – female tergum 8; 281 – female sternum 8. Scale bar 0.2 mm.

FIGURES 282 ­ 285. Aedeagus of Seeversiella luridicollis Gusarov, sp. n. (holotype). 282 – median lobe, parameral view; 283 – apex of median lobe, parameral view; 284 – median lobe, lateral view; 285 – apex of median lobe, lateral view. Scale bar 0.2 mm (282, 284), 0.1 mm (283, 285).

FIGURES 286 ­ 290. Genitalia of Seeversiella luridicollis Gusarov, sp. n. (286, 289, holotype; 287 ­ 288, 290, paratypes from Cerro Buenavista, Costa Rica). 286 – details of internal sac retracted into median lobe, lateral view; 287 – copulatory piece, lateral view; 288 – everted internal sac, parameral view; 289 – apex of left paramere; 290 ­ spermatheca. Scale bar 0.2 mm (286), 0.1 mm (287 ­ 290).

FIGURES 291 ­ 294. Abdominal segment 8 of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (291 ­ 292) and Cerro Buenavista (293 ­ 294), Costa Rica). 291 – male tergum 8; 292 – male sternum 8; 293 – female tergum 8; 294 – female sternum 8. Scale bar 0.2 mm.

FIGURES 295 ­ 298. Aedeagus of Seeversiella micralymma Gusarov, sp. n. (paratype from Cerro de la Muerte, Costa Rica). 295 – median lobe, parameral view; 296 – apex of median lobe, parameral view; 297 – median lobe, lateral view; 298 – apex of median lobe, lateral view. Scale bar 0.2 mm (295, 297), 0.1 mm (296, 298).

FIGURES 299 ­ 306. Genitalia of Seeversiella micralymma Gusarov, sp. n. (paratypes from Cerro de la Muerte (299, 301 ­ 305) and Cerro Buenavista (300, 306), Costa Rica). 299 – details of internal sac retracted into median lobe, lateral view; 300 – copulatory piece, lateral view; 301 – everted internal sac, lateral view; 302 – apex of left paramere; 303 – details of internal sac retracted into median lobe, abparameral view; 304 – everted internal sac, parameral view; 305 – medial lamellae, parameral view; 306 ­ spermatheca. Scale bar 0.2 mm (299, 301, 303 ­ 304), 0.1 mm (300, 302, 305 ­ 306).

FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 – male tergum 8; 308 – male sternum 8; 309 – female tergum 8; 310 – female sternum 8. Scale bar 0.2 mm.

FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320).

FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 – median lobe, parameral view; 322 – apex of median lobe, parameral view; 323 – median lobe, lateral view; 324 – apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324).

FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330).

FIGURES 331 ­ 334. Abdominal segment 8 of Seeversiella microphthalma Gusarov, sp. n. (331 ­ 332, holotype; 333 ­ 334, paratype from Villa Mills, Costa Rica). 331 – male tergum 8; 332 – male sternum 8; 333 – female tergum 8; 334 – female sternum 8. Scale bar 0.2 mm.

FIGURES 335 ­ 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 ­ 340, 342 ­ 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 ­ spermatheca; 342 ­ 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 ­ 343), 0.1 mm (336 ­ 337, 339, 341).

FIGURES 344 ­ 347. Abdominal segment 8 of Seeversiella geostiboides Gusarov, sp. n. (344 ­ 345, holotype; 346 ­ 347, paratype from Villa Mills, Costa Rica). 344 – male tergum 8; 345 – male sternum 8; 346 – female tergum 8; 347 – female sternum 8. Scale bar 0.2 mm.

FIGURES 348 ­ 355. Genitalia of Seeversiella geostiboides Gusarov, sp. n. (348 ­ 354, holotype; 355, paratype from Villa Mills, Costa Rica). 348 – median lobe, parameral view; 349 – apex of median lobe, parameral view; 350 – median lobe, lateral view; 351 – apex of median lobe, lateral view; 352 – apex of left paramere; 353 – details of internal sac retracted into median lobe, abparameral view; 354 – details of internal sac retracted into median lobe, lateral view; 355 ­ spermatheca. Scale bar 0.2 mm (348, 350, 353 ­ 354), 0.1 mm (349, 351 ­ 352, 355).

FIGURES 356 ­ 358. Abdominal segment 8 and spermatheca of Seeversiella adusta Gusarov, sp. n. (holotype). 356 – female tergum 8; 357 – female sternum 8; 358 ­ spermatheca. Scale bar 0.2 mm (356 ­ 357), 0.1 mm (358).

FIGURES 359 ­ 362. Abdominal segment 8 of Seeversiella flavida Gusarov, sp. n. (359 ­ 360, holotype; 361 ­ 362, paratype from Cerro Buenavista, Costa Rica). 359 – male tergum 8; 360 – male sternum 8; 361 – female tergum 8; 362 – female sternum 8. Scale bar 0.2 mm.

FIGURES 363 ­ 371. Genitalia of Seeversiella flavida Gusarov, sp. n. (363 ­ 367, 370, holotype; 368 ­ 369, 371, paratypes from Cerro Buenavista, Costa Rica). 363 – median lobe, parameral view; 364 – apex of median lobe, parameral view; 365 – median lobe, lateral view; 366 – apex of median lobe, lateral view; 367 – details of internal sac retracted into median lobe, lateral view; 368 – copulatory piece, lateral view; 369 – everted internal sac, parameral view; 370 – apex of left paramere; 371 ­ spermatheca. Scale bar 0.2 mm (363, 365, 367), 0.1 mm (364, 366, 368 ­ 371).

FIGURES 372 ­ 375. Abdominal segment 8 of Seeversiella paramoana Gusarov, sp. n. (372 ­ 373, holotype; 374 ­ 375, paratype from Cerro Chirripó, Costa Rica). 372 – male tergum 8; 373 – male sternum 8; 374 – female tergum 8; 375 – female sternum 8. Scale bar 0.2 mm.

FIGURES 376 ­ 384. Genitalia of Seeversiella paramoana Gusarov, sp. n. (376 ­ 380, 382 ­ 383, holotype; 381, 384, paratypes from Cerro Chirripó, Costa Rica). 376 – apex of left paramere; 377 – median lobe, parameral view; 378 – apex of median lobe, parameral view; 379 – median lobe, lateral view; 380 – apex of median lobe, lateral view; 381 ­ spermatheca; 382 – details of internal sac retracted into median lobe, abparameral view; 383 – details of internal sac retracted into median lobe, lateral view; 384 – everted internal sac, parameral view. Scale bar 0.1 mm (376, 378, 380 ­ 381, 384), 0.2 mm (377, 379, 382 ­ 383).