Tympanophorus borneensis, Rougemont, 2016

Rougemont, G. de, 2016, New Bornean Staphylinidae (Coleoptera), Linzer biologische Beiträge 48 (1), pp. 559-572 : 568-569

Tympanophorus borneensis nov.sp. (Fig. 7)

3 Holotype and 15 paratypes: MALAYSIA: Sabah, Lahad Datu, Ulu Segama For. Res., Danum Valley Forest Centre, 04º57.9’N 117º48.1’E, 200 m alt / xi.2005 1º For. baited PF, coll. Mann, Slade & Villanueva OUMNH-2006-051 / Tympanophorus N. Sp. Det. 2009 G. de Rougemont / Tympanophorus borneensis Des. 2015 G. de Rougemont ; 7 paratypes: MALAYSIA: Sabah, Lahad Datu, Ulu Segama For. Res., Danum Valley Forest Centre, 04º57.9’N 117º48.1’E, 450 m 20- 19.xi.2005 1º for. / Coll. Mann, Slade & Villanueva OUMNH2006-05 Lowland mixed dipterocarp for., West Trail area F.I.Trap ; 2 paratypes: MALAYSIA: Sabah, Lahad Datu, Ulu Segama For. Res., Coupe 81 logging area, 04º58.660’N 117º53.410E, iii.2005 FIT 2º Forest, Coll. E. Slade & J. Villanueva / Tympanophorus n. sp. det. G. de Rougemont (2011) / Slade colln. Ox. Uni. Mus. Nat. Hist. ( OUMNH) OUMNH-2005-062 ; 16 exx.: [same data as above localities] ; 1 paratype: SABAH, Danum Valley, B.R.L. f.i.t. 14-16.II.2007, G. de Rougemont [ CRO] ; 2 exx.: MALAYSIA, Sabah, Tawau , 11-17.x.2012, 4.66N 117.60E, 100 m alt., Coll. C.L. Gray / SAFE project area, F.I.T Riparian forest strip in oil palm, OUMNH-2013-056 .[Holotype, 17 paratypes and 18 other exx in OUMNH; 3 paratypes in CRO; 2 paratypes in LML] .

D e s c r i p t i o n: length: 10.5-11.5 mm. length of head: 1.3; breadth of head: 1.5; length of antenna: 2.5; length of pronotum: 2; maximum breadth of pronotum: 1.8; length of elytron: 2.2; breadth of elytra: 2.4. Body black, posterior 1/4 th of abdominal urite VIII reddish, antennae and legs pitchy. Vestiture consisting of mixed black and pale pubescence and black setae. Head transverse, eyes large, over three times longer than temples; puncturation rugose, homogenous. Antennomeres II-III obconical, of equal length, IV sub-quadrate, V-X increasingly transverse, Neck densely punctured, the punctures slightly transverse. Pronotum a little elongate, widest at anterior 1/3 rd the sides thereafter retracted to posterior angles; puncturation as coarse as that of head, in some individuals the interstices with a slight tendency to longitudinal confluence in middle; mid-line without any trace of impunctate areas. Scutellum strongly and densely punctured, the punctures transverse. Elytra sub-quadrate, the sides slightly dilated posteriad; sculpture coarser, denser and shallower than that of pronotum; antero-lateral parts of disc with sparse, decumbent, long pale setae in addition to short black pubescence. Abdominal tergites shiny, moderately densely punctures, the punctures rounded on tergite III, becoming strongly elongate on segments IV-VI, on tergite VII the punctures are large and elongate near anterior margin, becoming denser, round and finer posteriad, very fine near posterior margin.

M a l e: posterior margin of abdominal sternite VII (Fig. 7s7) slightly and shallowly concave; the sternite bears a small but conspicuous circular shiny impunctate impression just before posterior margin, the anterior rim of the impression with a pair of short stout hairs directed posteriad; sternite VIII strongly elongate, the posterior margin with a small rounded emargination; apex of sternite IX arcuately excised; aedoeagus; Fig. 7al; median lobe in ventral (Fig.7ml) view asymmetrical; paramere (Fig. 7p) extending beyond tip of median lobe, its apex with a small notch, and bearing an irregular cluster of ca. 15 peg setae and 8 fine pale apical and pre-apical setae.

In all males of this and other species of Tympanophorus that I have seen the inner sac of the aedoeagus is extruded, as it is in the illustrations of the two species described by SHIBATA (1975).

The large eyes of T. bornensis nov.sp. likens it to T. rugosus Waterhouse described from Timor, but the new species differs conspicuously in its smaller size and especially narrower build; in the new species the pronotum is elongate and the sides narrowed in posterior 2/3 rds whereas in rugosus the pronotum is a little transverse and widest in the middle, the elytra are smaller and less transverse, and the punctures of abdominal tergites much finer and sparser.The impression on the male 7 th sternite is much smaller than those of the only other species for which this character has been described, T. hayashidai SHIBATA , and T. sauteri BERNHAUER ( SHIBATA 1975) .