Themus (Telephorops) crassimargo Champion, 1926

Yang, Yuxia, Xi, Huacong, Yang, Xingke & Liu, Haoyu, 2019, Taxonomic review of the Themus (Telephorops) nepalensis species-group (Coleoptera, Cantharidae), ZooKeys 884, pp. 81-106 : 81

Themus (Telephorops) crassimargo Champion, 1926 Figs 3B , 4A, B , 9D

Themus crassimargo Champion, 1926: 127.

Themus (Tryblius) crassimargo : Wittmer 1975: 251, fig. 2 (aedeagus illustration).

Themus (Telephorops) subcaeruleiformis Wittmer, 1983b: 199, fig. 3, 3a (aedeagus illustration), syn. nov.

Themus (Telephorops) crassimargo : Okushima 1999: 59, figs 11 (habitus photo), 35 (female abdominal sternite VIII illustration).

Type material examined.

1♂ (BMNH, lectotype of Themus crassimargo ), [p]"Gopaldhara, Sikkim, vii.1924, H. Stevens", [p] “LECTOTYPUS”, [p] "Themus \ crassimargo \ Champ.". The lectotype was designated by Wittmer (1975).

1♂ (NHMB, holotype of Themus (Telephorops) subcaeruleiformis ), [h] "Kuanshien (Guanxian, now is Dujiangyan) \ Umg. VIII.1934", “600– 1300m \ Szechwan (Sichuan), China", [p] “HOLOTYPUS”, [h] "T. (Telephorops) subcaerulei- \ formis Wittm. \ det. W. Wittmer", [p] "Naturhist. \ Museum Basel \ coll. W. Wittmer", [p] "CANTHARIDAE \ CANTH00001233".

Other material examined.

CHINA, Xizang: 1♂ (CAUB), Zayü, Shajiong, 1700 m, 1978.VI.26, leg. F.S. Li. Sichuan: 1♀ (IZAS), 70 km West Chengdu, Qingcheng, Hou Shan mts., 1360 m, 30°44'N, 103°08'E, 2004.VIII.28, S. Murzin.

Distribution.

China (Xizang, Sichuan), N. India, Bhutan, Nepal.

Supplementary description.

Female ( Fig. 3B ). Like male, but antennomeres V–XI without impressions along outer edges (while present with smooth narrow longitudinal or oblong impressions in male), terminal abdominal ventrite wide (while narrow and triangular in male) ( Fig. 9D ) with posterior edge triangularly protuberant on each side, space between lateral protuberances about twice as wide as each width.

Remarks.

Themus (T.) subcaeruleiformis Wittmer, 1983 was originally described based on a single male type, from China, Szechwan, Kuanshien Umg. (now in Sichuan, Dujiangyan). Here a female ( Fig. 3B ) collected from Qingcheng, which is near the type locality, is discovered for the first time. The structure of its abdominal sternite VIII ( Fig. 9D ) is like that of T. crassimargo Champion, 1926, illustrated by Okushima (1999: fig. 35). Furthermore, T. subcaeruleiformis was only compared with T. subcaeruleus located in Yunnan, China in the original publication ( Wittmer 1983b), but not with species from the Himalayas ( Wittmer 1975).Moreover, the types of T. subcaeruleiformis and T. crassimargo were compared, but no differences between them were found in external morphology ( Fig. 4A, B ) and aedeagi illustrated by Wittmer (1975: fig. 2; 1983b: fig. 3, 3a).Therefore, T. subcaeruleiformis is proposed here to be junior synonym of T. crassimargo , according to the Principle of Priority ( ICZN 1999, Article 23.1).

Figures

Figure 3. Male habitus, dorsal view A Themus cavipennis Champion, 1926 (the specimen of Xizang) B T. crassimargo Champion, 1926 (the specimen of Sichuan). Scale bars: 5.0 mm.

Figure 4. Habitus, dorsal view A Themus crassimargo Champion, 1926 (lectotype) B Themus (Telephorops) subcaeruleiformis Wittmer, 1983 (holotype) C Triblius laboissierei Pic, 1929 (lectotype) D Themus (Tryblius) separandus Wittmer, 1975 (holotype).

Figure 9. Female abdominal sternite VIII, ventral view A Themus bicoloricornis Wittmer, 1983 B T. cavipennis Champion, 1926 C T. coelestis (Gorham, 1889) D T. crassimargo Champion, 1926 E T. crassipes Pic, 1929 F T. impressipennis (Fairmaire, 1886) G T. laboissierei (Pic, 1921) H T. masatakai Okushima, 2003 I T. sauteri (Pic, 1912) J T. uncinatus Wittmer, 1983. Scale bars: 1.0 mm.