Triphysa nervosa glacialis A. Bang-Haas, 1912

Dubatolov, Vladimir V., Korb, Stanislav K. & Yakovlev, Roman V., 2023, A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae), Ecologica Montenegrina 61, pp. 88-123 : 104-106

Triphysa nervosa glacialis A. Bang-Haas, 1912

Figs. 2m –x , 6a , 10–11 , 25 .

Triphysa dohrnii glacialis A. Bang-Haas, 1912 , Deutsch. Entomol. Zeit., Iris 26: 105.

Type locality: “Arasagun- Gol” [Arsan- Gol river , west environs of Khubsugul lake, Khubsugul Aymak, Mongolia] .

Type material (lectotype, designated here) in MHUB. Lectotype male, labelled: handwritten (black ink) on white paper “ Triphysa | Phryne | (Dohrni) | v. Glacialis | B.H”; printed on pink paper “Origin.”; printed on white paper “Arasagun- gol | Sajan”; printed on red paper: “ Lectotypus ♂ | Triphysa dohrnii | glacialis BangHaas 1912 | Dtsch. Ent. Z. Iris 26: 105 | Dubatolov, Korb, Yakovlev des. 2013”. Paralectotypes: 2 males, 4 females, same labels .

Diagnosis. In comparison to the nomenotypical subspecies the forewing upperside with cubital spots of the postdiscal series without white centers (as well as on hindwing), significantly larger than other postdiscal spots; the whole pattern is much mottled, veins on the forewing underside are contrasting. White edging of wings is very narrow. Fringes are brown.

Material examined. 1 male, 1 female, Altai ( ZISP) ; 1 female, Altai Max Bartel ( ZISP) ; 1 male, 1 female, S.E. Altai Tchuja Mts. 6-8000 ft. 5.7.[18]98 H.J. Elwes ( ZISP) ; 1 male, SE Altai, Tschuya Valley , 4- 6800 ft., 21-VI-1898, H. Elwes ( LNK) ; 2 males, S.E. Altai, Tshuya Valley , 4-6000f, 21-VI-[18]98, H.J.Elwes ( MHUB) ; 2 males, 2 females, Altai , Aktash, 7-12-VI-1992, S. Dialektov ( SCR) ; 1 male, Altai , Ulagansky distr., Yoldu river, 11-VI-2005, V.K.Zinchenko ( ISEA) ; 4 males, 4 females, S. Altai , Chagan-Uzun vill., 2300 m, 8-VII-1966, leg. A. Shtandel ( ZMMU) ; 5 males, 5 females, Altai , Kurai, 1800 m, 5-7- VI-1998, Aniskovitch leg. ( SCR) ; 5 males, 5 females, C. Altai , N. Chuya Mts., Maashei river., 1-4-VI-2001, I. Shiyanov leg. ( RYB) ; 1 male, N Ukok plateau, Tarkhata river valley, 2400 m, 27-VII-2013, S. Nikolaev ( ISEA) ; 1 male, 1 female, Ukok plateau, Zhumaly river upper course, 4-VII-2005, V.K. Zinchenko ( ISEAN) ; 3 males, 1 female, Ukok plateau, Muzdy-Bulak lake environs, 26-VI-1-VII-2005, V.K. Zinchenko ( ISEAN) ; 10 males, 10 females, SE Altai , Ukok plateau, Mai-pak, H – 2300 m, 29-VI-3-VII-1997, R. Yakovlev ( RYB) ; 1 male, 1 female, SE Altai, Ukok plateau, Mai-Tobe Mt., H – 2800 m, 7-VII-1997, R. Yakovlev ( RYB) ; 1 male, 1 female, SE Altai, Ukok plateau, Mai-Tobe Mt., H – 2800 m, 7-VII-1997, R. Yakovlev ( RYB) ; 1 male, 1 female, SE Altai , Ukok plateau, Bertek, H – 2200 m, 8-VII-1996, R. Yakovlev ( RYB) ; 1 male, 1 female, SE Altai , Kosh-Agatch, H – 1900 m, 23-VI-1993, E. Noskov ( RYB) ; 1 male, 1 female, SE Altai, Kosh-Agatch distr. , Dzhazaror (Belyashi), 21-VI-1995, H – 1800 m, R. Yakovlev ( RYB) ; 1 male, Kosh-Agach distr. , Zermokol-Nur lake, 2320 m, N49-56', E88-21', 22-VI-2005, V.K. Zinchenko ( ISEAN) ; 1 male, Kosh-Agach Distr. , upper Dzhazator River basin, Yuzhno-Chuiiskii Mt. Range , S slope, afternating dwarf, birch/ Kobresia tundras between the Akbul and Chikty Rivers, 2300-2500m alt. 15-VII-1998, O. Kosterin ( ISEAN) ; 1 male, 1 female, Altai , Kurai Mts., Cheibekkol’ lake, 20-25-V-1998, R. Yakovlev ( RYB) ; 3 males, 7 females, Kurai , Argatut valley, 6-20-VI-[19]60, Zheltikova ( ISEAN) ; 1 female, Kurai , Tyurgun’ valley, 20-VI-[19]60, Zheltikova ( ISEAN) ; 1 male, Sukor , near Chagan-Uzun, 2600 m, 7-VII- [19]66, A. Shtandel ( ISEAN) ; 1 male, 1 female, SE Altai, 15 km E Kokorya, Tapduair Mts. , 25-VI-2000, R. Yakovlev ( RYB) ; 2 males, Altai , Kurai, Tyurgun’ ( ISEAN) ; 2 males, Altai , Kurai, Argatut ( ISEAN) ; 1 male, near Chagan-Uzun ( ISEAN) ; 1 male, Altai , Krenrek, 29-VI-19…, leg. E. Rodd ( ISEAN) ; 1 male, Altai , North bank of Dzhulukul’ lake ( ISEAN) ; 1 male, Altai Nature Reserve , northern slope of Dzhulunkul lake, 2250 m, 2-VII-1986, N. Zolotukhin ( ISEAN) ; 1 female, Kultuk , Baicalsee ( LNK) ; 1 male, Baikal lake , Southern border of Baikalo-Lenskii Reserve ( ISEAN) ; 1 male, Arasagun gol, Sajan ( LNK) ; 7 males, 1 female Arasagun gol, Sayan (Co-types) ( ZFMK) ; 2 males, 2 females, Arasagun-gol Sajan ( ZISP) ; 3 males, 1 female, Sayany , Arasan-Gol, coll. Avinoff ( ZISP) ; 1 female, Kosogol [Khubsugul] lake, V. Dorogostajsky, 27.VI. ( ZISP) ; 1 male, 2 females, Sajan. mont ( ZISP) ; 1 male, 4 females, Gouv. Irkutsk , Sayan, orient Tunkinsk Weissberge, 2000 m, Juli ( ZFMK) ; 5 males, Munko Sardyk , Sayan Mts., Mondy-Irkutsk, 1800 m, August ( ZFMK) ; 1 male, Siberia or., Irkutzk, 25-V-1910, O. Hesse ( MHUB) ; 2 males, 2 females, E. Sayan, Mt. Khulugaisha , 25-VI-1979 ( ZSSM) ; 1 male, 1 female, Ost-Sajan ( MHUB) ; 1 male, 3 females, Sibiria or. ( MHUB) ; 1 male, Munko Sardyk , Sayan Mts., Mondy-Irkutsk, 1800 m, August ( MWM) ; 2 males, 1 female, Buryatiya , Mondy, 2-VII-1947, A. Tzvetaev ( ZMMU) ; 2 males, 2 females, Buryatiya , Mondy, 2000 m, 1- VI-2001 ( SCR) ; 1 male, 1 female, W. Buryariya, Mondy , 21-VI-2002 ( RYB) ; 2 males, 2 females, Buryatiya , Khamar-Daban, Taezhnyi, 10-VI-1988, Makhat leg. ( SCR) ; 2 males, 1 female, Buryatia , Kurumkan distr. , Dzherga Reserve, near Dzhirga, 27-V-1996, leg. S. Rudykh ( SRU) ; 1 male, Buryatia , Mondy, 14-VI-1986, Y.Timoshenko ( ISEAN) ; 1 male, Khakassiya , Shira lake, 15-VI-1931, P. Valdaev (MGU, Tzvetaev coll.) ; 2 males, 2 females, Minusinsk , Tagarskii Ostrov, 29-V-[19]33, Kozhantchikov ( ZSSM) ; 4 females, Shawyr , Tannu-Ola, Juni, 2500 m ( ZFMK) ; 1 male, Tuva , 6 km S Targalyk, 9-V-1990, V.K. Zinchenko ( ISEAN) ; 4 males, Tuva , Khovu-Aksy, Elegest river, 5-6-V-1990, V.K. Zinchenko ( ISEAN) ; 1 female, Tuva , Balagazinsky forest, 8-VI-[19]48 ( ISEAN) ; 2 males, Tuva , Khondergei, 22-6-[19]63, Violovich ( ISEAN) ; 2males, 2 females, Shawyr , Tannu-Ola, Juni, 2500 m ( ZSSM) ; 1 male, 1 female, Tuva , Ust-Khemsky distr., 25 km SSE Shagonar, Chelety river, 8-V-1990, V.K. Zinchenko ( ISEAN) ; 1 female, Tuva , 15 km S Shagonar, Chilekty, 10-V-1990, V.K. Zinchenko ( ISEAN) ; 1 male, 2 males, 2 females, Tuva , Kyzyl vic., right bank of Yenisey river, 20-V-1989, V.K. Zinchenko ( ISEAN) ; 2 males, Tuva , Kyzyl, mountains near upper course of Ulug-Khem river, 17-18-V-1990, V.V. Dubatolov, V.K. Zinchenko ( ISEAN) ; 1 male, 1 female, Tuva , Piy-Khema distr., Ust-Uyuk, 21-V-1989, V.K. Zinchenko ( ISEAN) ; 3 males, 3 females, W. Sayan, near Krasnoyarsk , Kacha river Valley, 15-V-2002 ( RYB) ; 1 female, Krasnoyarsk reg., near Solonzy vill., 12-VI-1977 ( ISEAN) ; 1 male, Krasnoyarsk reg., near Torgashino ( ISEAN) ; 1 male, Sayano- Shushenskii Reserve , river Uzun-Sug, 25-V-1989, leg. V. Vagin ( ISEAN) ; 1 male, 1 female, Sayano- Shushenskii Reserve , river Khem-Terek, 2-VI-1989, leg. V. Vagin ( ISEAN) ; 1 male, Irkutsk Prov. , southern border of Baikalo-Lensky Reservation, Onkholoi camp, 29-VII-2003, O. Berlov ; 1 female, Krasnoyarsk , Enisej Valley, A. Jacobson, 29.V.[8]97 ( ZISP) ; 3 males, 2 females, Torgashino , near Krasnoyarsk, A. Jacobson, 27.V.[18]97 ( ZISP) ; 1 male, 2 females, Krsnrsk [Krasnoyarsk], right bank, 29.V. ( ZISP) ; 1 male, 1 female, Minusinsk , Tag. [arsky] Island, 21.V.[19]24 ; 4 males, 1 female, 24.V.[19]24; 1 male, 6 females, 27.V.[19]24, Filipiev; 1 female, 19.V [19]24 ( ZISP) ; 1 male, Minusinsk , Tagarsky Island, 24.V.1930 ; 1 male, 28.V.1930; 1 male, 29.V.1930; 3 males, 30.V.1930; 3 males, 28.V.1931; 1 male, 2.VI.1931; 1 female, 22.V.1932, Kozhantschikoff; 3 males, 1 female, 23.V.1932; 2 males, 26.V.1932; 1 male, 28.V.1934 ( ZISP) ; 1 female, Togarsk. Isl. , 9.VI.1919 ( ZISP) ; 1 male, 1 female, near Minusinsk , 18.V.1916, Kozhantschikoff ; 1 male, 1 female, 19.V. ( ZISP) ; 1 female, Mishikha (Abakan Basin), Minus, A. Jacobson, 27.VI.[18]97 ( ZISP) ; 1 male, 3 females, Munko Sardyk Sajan mont ( ZISP) ; 1 male, Nukhu-Daban , 6.V.1914 S. Rodionoff ; 1 male, 26.V.1913 ( ZISP) ; 3 males, Shulun-Khurde , 29.V.1914, S. Rodionoff ( ZISP) ; 1 male, Irkutsk , coll. Avinoff ( ZISP) ; 1 female, Irkutsk ( ZISP) ; 3 males, 3 females, Irkutsk , 16 Mai [18]66, coll. Erschoff ( ZISP) ; 1 male, Irkut Leder кбвкНМ ( ZISP) ; 1 male, Irkut кбвкНМ ( ZISP) ; 1 male Irkutsk 4.VI.[19]09 A.I. Kiriloff ( ZISP) ; 1 male, Irkutsk , 15.V [19]07, Stschegolkova ( ZISP) ; 1 female, Irkutsk , 19/V [19]09, Stschegolkova ( ZISP) ; 2 female, Irkutsk ( ZISP) ; 1 female, Siberia Irkutsk coll.Meinhardt, 4101 ( ZISP) ; 1 male, 6 females, Verkhol. [enskaya] Mt. [near Irkutsk], 2/V [1]914 ; 16 males, 3/V [1]914, S. Rodionoff; 13 males, 2 females, 6.V ( ZISP) ; 1 male, 5 females, Glazun. Pad’ , 9.V.[1]914 S. Rodionoff ; 1 female, 7.V.1913; 1 male, 1 female, 27.V 1913 ( ZISP) ; 1 male, Gorkhon , 31.V 1914 S. Rodionoff ( ZISP) ; 1 male, Ushakovka , 12.V 1915, S. Rodionoff ( ZISP) ; 2 males, 4 females, Kachug river near Irkutsk, 15.V-15. VI Lukashik, Chiglinzev ( ZISP) ; 2 males, Irkusk gub. [Province] ( ZISP) ; 1 female, Petrovskij Zavod , Transbaikalia, 3-VI-[1]911, coll. Stschuko ( ZISP) .

Distribution. Russia: Russian Altai (Central and South-Eastern parts), Sayan Mountains (Krasnoyarsk reg., Khakasiya, Buryatiya near Mondy), Tuva (West and East Tannu-Ola Mts.), Khamar-Daban Mts., Irkutzk reg. (southern part), including western bank of Lake Baikal, Mongolia (Khubsugul distr.). Detailed distribution data in Altai and Sayan mountain regions listed by Yakovlev & Nakonechny (2001), Yakovlev (2004), Tshikolovets et al. (2009), in Buryatia – by Shodotova et al. (2007).

Figures

Figure 2. Triphysa sp., imago: a – T. striatula urumtshiensis, holotype (ZISP), upperside, b – underside; c – T. issykkulica sp. n., holotype (ZISP), upperside, d – underside; e – T. issykkulica sp. n., paratype, male, Kirgiziya, Issyk- Kul’ reg., Sarydzhaz, S slope of Inylchek Mts., Tashkoro, 3100 m, 4.07.1986, I. Pljustch (ISEAN), upperside, f – underside; g – T. issykkulica sp. n., paratype, female, 03.05.2009, Kazakhstan, W of Almaty, 800 m., leg. P.Egorov (SKNN), upperside, h – underside; i – T. issykkulica pljushchi ssp. n., holotype (ZMKU), upperside, j – underside; k – T. issykkulica pljushchi ssp. n., paratype (ZMKU), upperside, l – underside; m – T. nervosa glacialis, lectotype (MHUB), upperside, n – underside; o – T. nervosa glacialis, paralectotype, female (MHUB), upperside, p – underside; q – T. nervosa glacialis, male, SE Altai, Ukok plateau, Mai-pak, H – 2300 m, 29.06-3.07.1997, R. Yakovlev (RYB), upperside, r – underside; s – T. nervosa glacialis, female, SE Altai, Ukok plateau, Mai-pak, H – 2300 m, 29.06- 3.07.1997, R. Yakovlev (RYB), upperside, t – underside; u – T. nervosa glacialis, male, W. Buryariya, Mondy, 21.06.2002 (RYB), upperside, v – underside; w – T. nervosa glacialis, female, W. Buryariya, Mondy, 21.06.2002 (RYB), upperside, x – underside.

Figure 6. Distribution map of Triphysa species: a (above) – general map; b (below) – northern-eastern part of area of T. nervosa nervosa.

Figures 10−11. Habitats of Triphysa, and female of T. nervosa glacialis in nature: 10 – T. nervosa glacialis, SE Altai, Ukok plateau, near Bertek, h–2400 m (photo by R. Yakovlev); 11 − T. nervosa glacialis – SE Altai, near Chagan-Uzun vill., h–2200 m (photo by A. Volynkin).

Figures 25–30. Male genitalia (lateral projection) of Triphysa: 25 – T. nervosa glacialis, Altai, Ukok, Mai-Pak, h–2300 m; 26 – T. nervosa tuvinica, paratype; 27 – T. nervosa arturi, paratype; 28 – T. nervosa kobdoensis, paratype; 29 – T. nervosa mongolaltaica, holotype; 30 – T. nervosa biocellata, lectotype.