Glenoleon osmyloides (Gerstaecker, 1885)

Machado, Renato Jose Pires & Oswald, John David, 2020, Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae), Zootaxa 4796 (1), pp. 1-322 : 172-178

Glenoleon osmyloides (Gerstaecker, 1885)

( Figs. 85–87 )

Myrmeleon osmyloides Gerstaecker, 1885a:27 (OD); Gerstaecker 1885b:114 (cit).

Glenoleon osmyloides (Gerstaecker) : Esben-Petersen 1918:7 (ncb); Esben-Petersen 1923:578 (cit); Handschin 1935:690 (cit); Stange 1976:306 (cat); New 1985b:44 (key, rd, ill); New 1996:83 (cat); Stange 2004:105 (cat); Oswald 2018 (cat); Machado et al. 2019 (tree).

Glenoleon annulicornis Esben-Petersen, 1915:72 (OD) [as “ annulicorne ”]; Esben-Petersen 1917:205 (cit); Esben-Petersen 1918:7 (cit); Esben-Petersen 1923:578 (syn); Stange 1976:306 (cat); New 1985b:44 (cit); New 1996:83 (cat); Stange 2004:105 (cat); Oswald 2018 (cat).

Glenoleon gerstaeckeri New 1985b:42 (OD); New 1996:82 (cat); Stange 2004:104 (cat); Oswald 2018 (cat). new synonym

Diagnosis. Legs elongate; tibial spurs reaching T2 apex; forewing costal area uniareolate; hind wing without a broad band; forewing broad; male mediuncus forked apically in posterior view.

Description. Lengths: forewing: 19–24 mm; hind wing: 18–23 mm.

Head ( Figs. 85 a–b): Labrum and Clypeus pale and set with a line of elongate setae. Frons mostly black, except for ventrolateral corners pale (some specimens with the pale area little larger, taking part of ventral half); covered with short white setae. Gena pale. Vertex raised; mostly pale with two central and one posterior black marks in dorsal view (some species central marks fused); set with short black setae. Ocular setae absent. Antennae clubbed; elongate,> 2x length of pronotum; distance between antennae wider than scape width; apical flagellomeres dark brown, four to five subapical flagellomeres pale, remaining segments mostly light brown with pale areas, mainly on anterior surface of basal segments; torular membrane pale; flagellomeres almost as long as wide at base, but apical ones much wider than long; all segments set with short black setae. Mandibles pale with tip dark brown. Palpi, maxillary and labial pale, in some specimens the labial palpi segments are dark brown; apical labial palpomere fusiform, palpimacula opening oval-shaped, located medially.

Thorax ( Fig. 85b ): Pronotum longer than wide; posterior margin slightly wider than anterior; subapical furrow present; mostly pale with a central longitudinal black “Y” shaped line, and two longitudinal thin bands surrounding the central one (some darker specimens with black areas larger and fused); set with many short and long black setae, but with few long white setae at anterior margin. Mesonotum with prescutum and scutellum pale with a central black longitudinal band, scutum pale with black longitudinal bands (some darker specimens mostly black with small pale spots); covered with short black setae, and some elongate black setae, mainly on prescutum anteriorly. Metanotum with three longitudinal black bands, a central and two marginal, intercalated with pale bands; covered with short black setae. Pterothoracic pleura dorsal segments mostly black with scattered pale spots but ventral segments mostly pale with some black marks, mainly on dorsal margins (some darker specimens with black areas larger); covered with elongate white setae. Miller’s organ present.

Wings ( Fig. 85c ): Fairly broad with tip acute; anterior Banksian line present in both wings, posterior absent; veins set with short black setae, in general brown, except by the main longitudinal veins that are intercalated by numerous short white regions. Male pilula axillaris present. Forewing membrane almost hyaline with small brown marks as follows: on posterior margin, tip, pterostigma, subcostal area, around mediocubital crossveins, one small brown line on prefork area apex, and another on rhegmal area; CuA fork located between RP origin and first fork; three or four presectoral crossveins (specimens with four the distal one bifurcates); costal area large; with many subcostal veinlets forked; posterior area narrower than prefork area. Hind wing membrane hyaline except for a small brown mark on rhegmal area; MP fork located between RP origin and first fork; subcostal veinlets simple but few distal ones forked in some specimens; one presectoral crossvein.

Legs ( Figs. 85 a–b): All pairs of legs, femur elongate (> 3.5x length of coxa); tibia about same size of femur but longer than tarsi; tibial spurs surpassing T2 tip; T2, T3 and T4 about same size, T1 about twice as long as T2, T5 about twice as long as T1; claws about half of T5 length; coxa, trochanter, and femur set with short white setae; femur, tibia and tarsi set with short black setae, and scattered long black setae; T5 ventrally with two rows of thick, long, black setae. Proleg femur sense hair absent; tibia with antennal cleaning setae ventroapically; coxa mostly pale with irregular black marks; trochanter pale; femur pale with tip dark brown; tibia mostly pale with three dark brown rings; tarsi with T3, T4 and tip of T5 dark brown, remaining areas pale. Mesoleg with same colour pattern of proleg. Metaleg with femur, tibia and T1 slightly longer than in other legs; colouration similar to other legs, except for tibia pale, with tip dark brown.

Abdomen: In general, mostly black with many irregular pale marks, but in some specimens the pale marks are dominant. Beset with short black setae.

Male Terminalia ( Figs. 86 a–d): Ectoproct posterior margin rounded in lateral view, set with elongate black setae. 9 th sternite in ventral view, with large medial invagination; covered with elongate black setae. Gonarcus arched with lateral areas broad in posterior view; curved in lateral view. Mediuncus short and forked at tip. Paramere broad with an acute medial extension in lateral view.

Female Terminalia ( Figs. 86 e–f): Ectoproct posterior margin rounded and covered with thin elongate setae. Lateral gonapophyses rounded, smaller than ectoproct, set with cavisetae. 7 th sternite distal margin straight in ventral view; covered with long black setae. Pregenital plate elongate longitudinally, with posterior margin rounded in ventral view. Posterior gonapophyses broad, elongate, with tip slightly enlarged; covered with long black setae, and cavisetae on apex. 9 th tergite with an elongate membranous digitiform process. Anterior gonapophyses as two small rounded lobes, set with few thickened setae. Ventral membrane, gonapophyseal plates present.

Distribution ( Fig. 87 ). Australia: NSW, NT, QLD, SA, VIC, WA. Widespread and common across mainland

Australia.

Adult activity period. Records for all months of the year; most records for October to February.

Biology. Unknown, larva unknown.

Name-bearing types. Myrmeleon osmyloides : Holotype (by implicit monotypy), male [not female], ZIMG, high-resolution images examined. Type locality: Australia. From original description: “(Fem.).—Patria: Australia ”. From New (1985b): “Types / Holotype of Myrmeleon osmyloides 'Australia' (Greifswald, not seen)”. From New (1996): “Type data: holotype EMAG adult sex indet.*. / Type locality: Australia ”.

Gerstaecker’s (1885a) statement that the apparently single specimen upon which the species was based is a female is incorrect; the presence of a pilula axillaris at the base of the hind wing demonstrates that the holotype is a male. Condition: good, missing left hind wing and hind leg.

Glenoleon annulicornis : Holotype (by original designation), female, BMNH, high resolution images examined. Type locality: Australia. From original description: “ Victoria; one specimen (Coll. Esben-Petersen), the other not labeled. Type in Coll. West Australian Museum, Perth; cotype in Coll. Esben-Petersen”. From New (1985b): “ G. annulicorne was described from two specimens: the type, designated as having no data, stated to be in WAM. I did not find any such specimen in WAM. There is in BMNH, though, a specimen of this species, tallying exactly in leg positions with Esben-Petersen's photograph, and also lacking abdomen (also now antennae): labelled as Type (seen). I believe this to be the type”. From ( New 1996): “Type data: syntypes BMNH 1 adult sex indet. (whereabouts of other syntypes unknown)”.

The statements made by New (1985b) are confirmed here; the holotype stated by Esben-Petersen (as “Type”)

to be deposited in the WAM is now in the BMNH. The specimen is identical to the picture presented in the original description and also bears a type label. The sex of the holotype was never mentioned but it is clearly a female because of the absence of the pilula axillaris. The “cotype” (a paratype) with location unknown was located during this study. It is a male deposited in the ZMUC; labels: Vict. (small pale square), Glenoleon annulicorne n. sp. Esben Petersen Cotype (large white rectangle), cotype (elongate red rectangle). Specimen in general well preserved: right meso leg missing, and tip of the wings damaged.

Glenoleon gerstaeckeri : Holotype (by original designation), male, AMSA, high-resolution images examined. Type locality: Australia, Western Australia, Kalbarrie (town). From original description: “ Holotype, ♂, Western Australia, Kalbarrie, 22.xi.1978, M.S. and B.J. Moulds (AM)”. Condition: good; right metaleg missing; terminalia dissected.

Additional material examined (97♂, 172♀, 13?). AUSTRALIA: ACT: Canberra : 16.ix.1957, E. F. Riek (1♀, FSCA) ; 4.i.1931, T. Greaves (1♂, ANIC) ; 16.xii.1958, E.F. Riek (1♀, ANIC) ; 20.ii.1960 (1♀, ANIC) ; 1.iv.1957 (1♀, ANIC) ; 15.xii.1960 (1♀, ANIC) ; 19.xi.1959 (1♀, ANIC) ; 12.i.1960 (1♂, ANIC) ; 4.i.1960 (1♂, ANIC) ; 8.ii.1961 (1♀, 1♂, FSCA) ; New South Wales: Albury : xii.1960 (1♀, FSCA) ; Bourke : 27.x.1949, E.F. Riek (1♀, 1♂, ANIC) ; 19.xi.1949 (3♀, 1♂, ANIC) ; Broken Hill : near race track , 280m, 31.96445 oS– 141.41817 oE ± 70m, 2.i.2013, M.V., Oswald, Diehl & Machado, Oswald #625 (1♀, TAMU); Coll. Lower (1♀, ANIC) ; 20.xi.1949, E.F. Riek (10♀, ANIC) ; 19.xi.1949 (7♀, 2♂, ANIC) ; vi.1946 (1♀, ANIC) ; Canowindra : 1.i.1955 (1♂, MVM) ; 30.xii.1954 (1♀, MVM) ; 70 W of Cobar , 24.xi.1949 (3♀, 2♂, ANIC) ; Dareton : 29.xi.1993, N.W. Rodd (1♂, AMSA) ; Dowsons Springs: Mount Kaputar , 18.ix.1968, C.W. Frazier (1♀, 1♂, ANIC) ; 30.x.1967 (3♀, 3♂, ANIC) ; 16 km of Eubalong West , 27.xii.1992, M.S. & B.J. Moulds (1♀, AMSA) ; Hankes : x.1966 (1♀ SAMA) ; Grassmere Station : ix.1962 (1♂, SAMA) ; Nombinnie Nature Reserve: 18 air km SSE Mount Hope , 160m, 32.99613 oS– 145.94006 oE ± 70m, 3.i.2013, M.V., Oswald, Diehl & Machado, Oswald #626 (1♂, TAMU) ; Nyngan : 21.x.1949, E.F. Riek (3♀, ANIC) ; Pilliga Nature Reserve : 34 km N Coonabarabran, 23.ix.2005, R. Miller & L. Stange (1♂, FSCA) ; Trangie : 13.vi.1968 (1♀, ANIC) ; Yanco : 8.xi.1949 (3♀, 1♂, ANIC) ; Yass : 26.xii.1931 (1♀, ANIC) ; Walgett : 15.x.1957, E.F. Riek (1♀, ANIC) ; Wilcannia : 22.xi.1949, E.F. Riek (1♀, ANIC) ; Northern Territory: Alice Springs : 19.x.1966 (1♀, ANIC) ; 17.ii.1966 (1♂, FSCA) ; x.1978 (1♀, ANIC) ; Austral Downs : ix.1977 (1♂, 2♀ SAMA) ; Bullita homestead: 21.vi.1969 (1♀, ANIC) ; Clifton Hills : 18.ix.1972, M.S. Upton (3♀, 4♂, ANIC) ; Coolibah : 15.34S– 130.54E, 15.ix.1968 (3♀, ANIC) ; 10.vi.1968 (3♂, ANIC) ; Erldunda : ix.1967 (1♀, MVM) ; Hermansberg : iii.1967 (1♂, SAMA) ; Katherine : viii.1973 (2♀, 2♂, ANIC) ; Kings Canyon : 22.xi.1989, R. Miller (1♂, 1♀, FSCA) ; Lake Woods : x.1977 (2♀, SAMA) ; Limbunya : 2.viii.1969 (1♀, ANIC) ; Newcastle Waters : 4.vi.1929, T.G. Campbell (1♂, ANIC) ; Victoria River : 18 km W of Timber Creek township, 25.xii.1991, M.S. & B.J. Moulds (1♀, AMSA) ; ix.1973 (1♀, ANIC) ; Yuenduni : ii.1968 (1♂, SAMA) ; Queensland: Archer River crossing: N of Coen, 7.x.1979, M.S. & B.J. Moulds (1♀, AMSA) ; Atherton Tableland : x.1947, N. Geary (1♀, AMSA) ; Barakula State Forest : 28.5 air km NNW Chin- chilla, 26.49661°S 150.52354°E, R Machado, MV light, 19.xii.2015, AustRM#04 (1♂, TAMU) ; Bingil Bay : 30.vi.1992, @ light, DH Habeck (1♂, FSCA) ; Bollon : 35 N, 17.x.1957, E.F. Riek (1♂, 1♀, ANIC) ; Boulia : x.1978 (1♀, ANIC) ; Brisbane : (9?, QMBA) ; 23.x.1926 (1♂, ZMUC; 1♀, QMBA); Comooweal : 13.vi.1968 (2♀, ANIC) ; Carandotta : ix.1977 (3♂, 1♀ SAMA) ; Charleville : 11.i.1934 (1♀, QMBA) ; 21.x.1957 (5♀, 1♂, ANIC) ; x.1975 (1♀, ANIC) ; Chinchilla : 20.x.1985 (1♀, QMBA) ; 1.i.1986 (1♀, QMBA) ; Clermont : Dr. K.K. Spence (1♂, 1♀, AMSA) ; Cunnamulla : 20.x.1938, N. Geary (1♂, AMSA) ; x.1941, N. Geary (1♂, AMSA) ; x.1943, N. Geary (1♀, AMSA) ; ii.1949, N. Geary (1♀, AMSA) ; Durak River crossing: Gibb River road, E. Kimberley, 29.xii.1991, M.S. & B.J. Moulds (1♂, AMSA) ; Eidsvold : (1♀, ANIC) ; 6.x.1936 (1?, QMBA) ; Emerald : 20.ii.1916 (1♀, 1♂, ANIC) ; Eromanga : xi.1967 (1♀, ANIC) ; Gregory River: at Gregory Downs , 20.xii.1986, M.S. & B.J. Moulds (1♀, AMSA) ; Julia Creek : 18.xii.1954 (1♀, QMBA) ; Kundala : (1♀, MVM) ; Lawn Hill National Park : 3.v.1995 (1♀, QMBA) ; Longreach : 11.xi.2005, L. Stange (1♀, FSCA) ; Magnetic Island : (1♀, SAMA) ; Mitchell : 18.x.1957, E.F. Riek (1♀, ANIC) ; Mount Isa : i.1992 (1♀, ANIC) ; Muttaburra : 22.viii.1966, G. Monteith (1♂, QMBA) ; Nappamerry : 6.xi.1949, E.F. Riek (2♀, ANIC) ; Nockatunga : 10.xi.1949, E.F. Riek (1♀, ANIC) ; Powela Aramac : viii.1920, F. Bradshaw (1♀, AMSA) ; Roma : 11.xi.1938 (2?, QMBA) ; Silver Plains : 16.xi.1996 (1♀, ANIC) ; Sunnybank : 18.x.1926 (1♂, QMBA) ; 28 km SE of Tambo , 11.ii.1981, M.S. & B.J. Moulds (1♀, AMSA) ; Thargomindah : 22.x.1957, E.F. Riek (1♀, ANIC) ; Taroom District: Boggomoss 21, 25 o 27’S– 150 o 03’E, 10.xi.1996, at light, C.J. Burwell & S. Evans (1♀, QMBA) ; Yulepa : (1?, QMBA) ; Walkers Creek : 35 km NNE of Normanton, 2.i.1990, M.S. & B.J. Moulds (1♂, AMSA) ; Westwood : 29.ix.1923 (1♀, MVM) ; Windorah : 3.ix.1997 (1♀, QMBA) ; South Australia: Adelaide : xii.1886 (1♀, SAMA) ; Andamooka Homestead : x.1975 (1♂, SAMA) ; Approdinna Attora Knolls : ix.1998 (1♀, SAMA) ; Arkaba Creek: 26 air km ENNE Hawker , 390m, 31.68717 oS– 138.57257 oE ± 70m, 31.xii.2012, M.V., Oswald, Diehl & Machado, Oswald #623 (1♀, TAMU) ; Flinders Range : iii.1973 (1♀ SAMA) ; Arkaroola: northern Flinders Ranges [Vulkathunha-Gammon Ranges Park], 22.i.1976, M.S. & B.J. Moulds (1♂, AMSA) ; xi.1969 (2♂, 1♀, SAMA) ; Athelstone : x.1973 (1♀, SAMA) ; Balcanoona Creek : xi.1975 (1♀, SAMA) ; Billero Dam : xi.1975 (3♀, SAMA) ; Copley : x.1967 (1♂, 1♀, SAMA) ; Coward Springs : xi.1955 (1♀, SAMA) ; Dalhousie Springs : x.1987 (1♀, SAMA) ; x.1980 (1♂, SAMA) ; Finniss Springs Homestead : xii.1964 (1♂, SAMA) ; Gawler Ranges National Park : x.2006 (1♂, SAMA) ; Gluepot Reserve : xii.2000 (1♂, SAMA) ; Goyder Lagoon : 18.ix.1972 (4♂, ANIC) ; Hesso : x.1975 (1♂, SAMA) ; Indulkana Creek : ix.2001 (1♀, SAMA) ; Innamincka: Cooper Creek , 24.i.1976, M.S. & B.J. Moulds (1♀, AMSA) ; ix.1996 (1♀, SAMA) ; Kalamurina Homestead : 17.ix.1972 (1♀, ANIC) ; Karoonda : (1♀, SAMA) ; Koonchera : 2.ix.1997 (2♀, QMBA) ; Lake Eyre: Prescott Point , iii.1965 (2♂, SAMA) ; Leigh Creek : 14.xii.1966 (1♂, ANIC) ; x.1965 (1♂, ANIC) ; xii.1965 (2♂, 5♀ SAMA) ; Marree : 1914 (1♀, ANIC) ; 15.ix.1972, Z. Liepa (4♂, ANIC) ; xii.1964 (1♀, SAMA) ; x.1966 (1♂, 1♀, SAMA) ; McDonalds Downs : viii.1930 (2♂, 1♀, SAMA) ; Mitchan : ii.1975 (1♀ SAMA) ; xii.1978 (1♀ SAMA) ; Mount Freeling Station : xi.1998 (2♂, SAMA) ; Mulloorina Homestead : iii.1965 (1♀ SAMA) ; Munyaroo Conservation Park : xi.2005 (1♀, SAMA) ; Murta : x.1960 (1♀, SAMA) ; Nullarbor National Park : x.2004 (1♀, SAMA) ; Oraparinna Creek : 32 o 21’S– 138 o 42’E, 9.x.1997, J. & A. Skevington, C. Lambkin, S. Winterton (1♀, QMBA) ; Reynella : x.1922 (1♂, SAMA) ; Yelpawaralinna : ix.1993 (2♀, SAMA) ; White Bull Yard: Kalamurina Station x.1999 (1♂, 1♀, SAMA) ; Wirreanda Creek : xi.1975 (1♂, SAMA) ; Victoria: (1♀, ANIC) ; Deddick River Junction: Snowy River , 14.xii.1976 (1♀, MVM) ; Inglewood : 1902 (1♂, ANIC) ; Kewell : iv.1988 (1♂, ANIC) ; Kyabram : 1947 (1♂, MVM) ; 25.xi.1945 (1♀, MVM) ; Mildura : 9.xi.1947 (3♀, MVM) ; Ouyen: Mallee District (1♂, MVM) ; Western Australia: (1♀, FSCA) ; Carnarvon : x.1992 (1♂, ANIC) ; Cave Hill : iii.1996 (1♂, SAMA) ; Claremont : i.1914 (1♀, ANIC) ; Dampier : 20.iv.1971 (1♀, ANIC) ; Drys- dale River National Park: Carson River St. , 14 o 37’S– 126 o 56’E, 1.viii.1996, Edwards (1♀, MVM) ; Halls Creek : iii.1995 (1♀, ANIC) ; Hancock Range : v.2003 (1♀, ANIC) ; Kalgoorlie : 30.xi.1985, M.S. & B.J. Moulds (2♀, AMSA) ; 20.i.1989, A.J. Graham (1♂, AMSA) ; Karijini Drive: 82 km S junction on Great Northern highway, 694m, 23 o 07.3’S– 119 o 05.5’E, Malaise trap in wash with pools, 23.iv–16.v.2003, M.E. Irwin & F.D. Parker (1♂, FSCA) ; Karijini National Park: Kalamina Road , 648m, 22 o 25.6’S– 118 o 23.7’E, Malaise trap across ravine leading to gorge, 23.iv–4.v.2003, M.E. Irwin & F.D. Parker (3♀, FSCA) ; Karonie : 9.xi.1969 (2♀, ANIC) ; Kelmscott : 19.i.1936, K.R. Norris (1♂, ANIC) ; Kunamurra : ii.1979 (1♀, ANIC) ; iv.1966 (1♀ SAMA) ; Mandora station : ix.1979 (1♀, ANIC) ; Marble Bar : 18.ii.1977, M.S. & B.J. Moulds (1♀, AMSA) ; 30mi S of Mendurah , 13.i.1971, mv lamp, G.A. Hollo- way & H. Hughes (1♂, 1♀, AMSA) ; Millstream Homestead : 2.iv.1971 (1♂, ANIC) ; 2.xi.1970 (2♀, ANIC) ; Ned- lands: iii.1963 (1♂, SAMA) ; Norseman : 17.xi.1969, Key & Upton (3♀, ANIC) ; Serpentine Falls: Darling Ranges , 30.xii.1970, mv lamp, G.A. Holloway & H. Hughes (1♂, AMSA) ; 20.xi.1978 (1♂, MVM) ; Tickalara : 14.xi.1949 (1♂, ANIC) ; Wandoo National Park: Kent Road, 26 km air SWSW York , 270m; 31.99773 oS– 116.52801 oE ± 70m, 17.xii.2012, M.V., Oswald, Diehl & Machado, Oswald #606 (1♀, TAMU) ; Weeli Wolli Creek: Hancock Range, 610m, Malaise trap, 26 km air SWSW York , 270m; 23 o 1.9’S– 119 o 10.7’, 6–17.v.2003, M.I. Irwin & F.D. Parker (2♀, FSCA) ; Widgiemooltha : 31.14S– 121.28E, 6.xi.1969, Key & Upton (1♀, 1♂, ANIC) ; Wyndham : 15.vii.1969 (1♂, MVM) ; no label: (1♂, ANIC) .

Comments. Synonymy of Glenoleon annulicornis: The synonymy of Glenoleon annulicornis under G. osmyloides , first proposed by Esben-Petersen (1923) and later corroborated by New (1985b), is confirmed here once again. The two specimens of the type series of G. annulicornis were re-examined and it is clear that both belong to G. osmyloides .

Synonymy of Glenoleon gerstaeckeri: In the original description of G. gerstaeckeri New (1985b) noted its resemblance to G. osmyloides , but differentiated the species based on the abdominal pattern and genitalia. Re-examination of the holotype of G. gerstaeckeri for this work has shown it to be very similar to G. osmyloides -- its general body marks, leg lengths, and wing shape and venation are almost identical. The abdominal pattern mentioned by New is not a robust character and is one of the most variable characteristics of the Periclystus genus group. The holotype is only slightly paler than typical G. osmyloides specimens. The genitalia are essentially identical to G. osmyloides , the only difference being the slightly wider parameres (a character too weak to sustain the species). Based on these observations Glenoleon gerstaeckeri is here synonymized with Glenoleon osmyloides .

Figures

FIGURE 85. Glenoleon osmyloides: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal.

FIGURE 86. Glenoleon osmyloides: male: a) terminalia, lateral; b) genitalia, lateral; c) genitalia, posterodorsal; d) 9th sternite and genitalia, ventral; female: e) terminalia, lateral; f) terminalia, ventral (note pregenital plate hidden beneath sternite). Scale bars = 0.5 mm.

FIGURE 87. Distribution of Glenoleon species (part).