Chelipoda manggawna, Plant, 2009

Plant, Adrian R., 2009, Diversity Of Chelipoda Macquart, 1823 (Diptera: Empididae: Hemerodromiinae) In Northern Thailand With Discussion Of A Biodiversity ‘ Hot Spot’ At Doi Inthanon, Raffles Bulletin of Zoology 57 (2), pp. 255-277 : 266-267

Chelipoda manggawna , new species

( Figs. 10 , 25 )

Material examined. – Holotype. Male, THAILAND: Chiang Mai, Doi Inthanon National Park, Checkpoint 2, 18°31.554'N 98°29.940'E, 1,700 m, Malaise trap, coll. Y. Areeluck, 16–23 Mar.2007 ( QSBG, T1817 ).

Paratypes. Same data as holotype : 1 male, 30 Aug. –6 Sep.2006 ( NMWC, T238 ) ; 1 male, 6–13 Sep.2006 ( QSBG, T244 ) ; 1 male, 1 female, 13–21 Sep.2006 ( NMWC, T250 ) ; 1 female, 19–26 Oct.2006 ( NMWC, T377 ) ; 2 females, 2–10 Nov.2006 ( QSBG, T389 ) ; 1 male, 17–24 Nov.2006 ( QSBG, T1846 ) ; 4 females, 24 Nov. –1 Dec.2006 ( NMWC, T1870 ) ; 1 male, 2–9 Feb.2007 ( QSBG, T1793 ) ; 1 female, 16–23 Feb.2007 ( QSBG, T1805 ) ; 1 male, 23 Feb. –2 Mar.2007 ( NMWC, T1775 ) ; 3 males, 16–23 Mar.2007 ( QSBG, T1817 ) ; 3 males, 16–23 Mar.2007 ( NMWC, T1817 ) ; 2 females, 15–22 Apr.2007 ( QSBG, T1845 ) ; 1 female, 29 Apr. –6 May.2007 ( QSBG, T1857 ) ; 1 male, 1 female 1–8 May.2007 ( NMWC, T1827 ) : Kew Mae Pan Trail, 18°33.162’N, 98°28.810’E, 2,200 m, 1 female, 1–8 May.2007 ( NMWC, T1824 ) ; 1 female, 2–9 Mar.2007 ( QSBG, T1777 ) : Campground Pond, 18°32.40'N 98°31.80'E, 1,200 m, 1 male, 5–12 Jan.2007 ( QSBG, T1917 ) : Summit Marsh, 18°35.361'N 98°29.157'E, 2,500 m, pan trap, 1 male 28 Feb.2007 ( NMWC, T1767 ) .

Diagnosis. – Yellow species without dark median stripe on scutum and legs somewhat stouter than in C. flavida . Head yellow, only ocellar protuberance black.

Description. – Male. Body length 3.0mm. Head yellow, ocellar protuberance black. Setae blackish, strong, including one ocl, one vtl and one upper upo; one or two minute upo below upper upo but occiput otherwise lacking upo and lpo and only a few minute setulae behind mouth. Antenna with basal segments yellow; poped 2× long as wide, dark brown; stylus 4× long as poped, dark brown to yellowish brown. Mouthparts yellow.

Thorax. Dorsum yellow, pleura pale yellow, lateral margins of scutum narrowly darkened. Setae blackish; anterior dc and upper npl strong, posterior dc (in line with npl) much weaker and of similar size to sa and sct. Lower npl much smaller, anterior postpronotal setulae minute with 2–4 rather longer setulae between it and notopleural area.

Legs yellow, tarsomeres 4–5 darker. C 1 as long as thorax ( Fig. 25 ), rather slender, 9.5× as long as wide, anterobasal setae not distinguished from regular row of yellowish anterior setulae. F 1 as long as C 1, distinctly inflated, 4.6× as long as wide, widest 0.4–0.5 from base; femoral formula 5(4–5)/23(18–25)/13(11–16)/4(3–4) +1 ( Table 1), spines yellowish, denticles black, pv spines very weak distally; basal spine rather weak, more or less contiguous with series of av spines.T 1 0.65× long as F 1.

Abdomen brownish yellow, paler ventrally; tergite 8 reduced; sternite 8 with distinct but short setae. Epan and Hypan ( Fig. 10 ) similar to C. flavida , fused, erect, narrowly subrectangular in lateral view, yellowish, only small setae posteriorly, left and right lamellae distinctly separated by unpigmented membrane. Cercus fused with Epan; a short digitiform yellow process bearing strong yellow setae apically, apparently emerging from the inner face of a somewhat darker rounded and rather undifferentiated basal lobe which bears more numerous smaller setae. Subepandrial process blackish, short and robust, conspicuously bent at 0.5 from base, flattened above apically but convex below. Postgonite narrow, apically upturned, black, basally yellowish and broader. Phallus narrow, yellow, slightly upturned apically, reaching as far as apex of cercus.

Wing membrane faintly brownish, veins brown. Squamae with dark fringes. Halter greyish white, stems darker.

Female. Head with setae rather stronger than male. F 1 slightly more inflated, 4.3× long as wide, widest 0.3 from base; femoral formula 5(5–6)/24(21–27)/14(11–16)/4(4–5) +1 ( Table 1); av and pv spines blackish or dark yellow. Abdomen brownish dorsally, whitish yellow ventrally; sternite 8 dark brown, subquadrate. Cercus elongate, yellow.

Etymology. – The specific epithet derives from the Thai word for dragon, mang-gawn; a reference to supposed fierce predatory behaviour inferred by the presence of raptorial front legs.

Remarks. – This species is known only from Thailand’s Doi Inthanon between August and May from 1,200 –2,500 m (mostly 1,700 m).

Figures

Figs. 7–11. Male genitalia of Chelipoda species in lateral view: 7. C. kameawuta new species; 8. C. laisoma new species; 9. C. macrosceles new species; 10. C. manggawna new species; 11. C. meenamluang new species.

Figs. 25–28. Chelipoda species: 25. C. manggawna new species, male; 26. C. macrosceles new species, female habitus; 27. C. nakropa new species, male, front femora showing chaetotaxy; 28. C. hubeiensis Yang & Yang, female, wing. Abbreviations: bm, cell bm; br, cell br; bs, basal spines; d, denticles; dm, cell dm; dm-cu, crossvein dm-cu; s, spines.