Drilus bicolor Schaufuss, 1867

Kundrata, Robin, Kobieluszova, Lucie & Bocak, Ladislav, 2014, A review of Drilini (Coleoptera: Elateridae: Agrypninae) of the Northern Levant, with description of a new species from Syria and a key to Levantine species, Zootaxa 3755 (5), pp. 457-469 : 463-464

Drilus bicolor Schaufuss, 1867

( Figs 3 , 9, 15 , 21 )

Drilus bicolor Schaufuss, 1867: 83 .

Material examined. 1 male, SYRIE, PIC 1899 // Beyruth ( MHNP).

Diagnosis. Drilus bicolor is easily recognizable from its Levantine congeners by bicolored elytra, which are lighter only at humeri ( Fig. 3 ), shorter antennomere 3 ( Fig. 9 ), and pronotum with fine, deep puncturation, and acutely projected hind angles ( Fig. 15 ).

Redescription. Male. Body medium-sized, elongate, 3.1 times as long as width at humeri, moderately convex dorsally ( Fig. 3 ). All body parts yellowish to brown, only elytral apical four fifths dark brown; entire body covered with yellow pubescence.

Head slightly hypognathous, including eyes as wide as anterior margin of pronotum. Surface of cranium punctured, wrinkled, dorsally with long sparse semierect pubescence. Eyes medium-sized, hemispherically prominent, their frontal distance 2.1 times eye diameter. Mandibles robust, long, considerably curved, incisor margin with conspicuous tooth in middle part, shiny. Maxillary palpi slender; apical palpomere obliquely cut; labial palpi tiny, apical palpomere pointed. Antennae 11-segmented, reaching third of elytral length, scapus robust, pedicel short, small, antennomere 3 serrate, about 2.5 times longer than antennomere 2, antennomeres 4–10 shortly flabellate, subequal in length, apical antennomere simple, longest, slightly longer than lamella of penultimate antennomere ( Fig. 9 ).

Pronotum flat, transverse, widest at basal margin, 1.7 times wider posteriorly than length at midline. Anterior margin straight, lateral margins convex, posterior margin slightly sinuate in middle. Anterior angles almost rectangular, posterior angles considerably prominent, acutely projected ( Fig. 15 ); surface of disc with fine punctures, distributed sparsely in middle and densely at margins, with sparse, semierect, long setae. Scutellum flat, triangle-shaped. Prosternum transverse, prosternal process short, slightly elevated in middle part. Mesoventrite narrow, v-shaped. Metaventrite large, trapezoidal, sparsely punctured. Elytra subparallel-sided, slightly widened between humeri and half of elytral length, 2.5 times longer than width at humeri, tapered apically, covered with sparse, semierect pubescence. Abdomen slender, ventrites with fine microstructure and sparse, long hairs. Legs slender, slightly compressed, with sparse, long, semierect setae, coxae long, robust, trochanters slender, obliquely attached to femora, tarsomeres 1–3 subequal in length, tarsomere 4 shortest, apical tarsomere long, claws simple, slightly curved.

Male genitalia with robust phallus; phallus longer than paramerae, considerably curved; short robust paramerae with sparse setae; phallobase long ( Fig. 21 ).

Measurements. BL 5.4 mm, EL 4.3 mm, WH 1.7 mm, PL 0.8 mm, PW 1.4 mm, Edist 0.7 mm, Ediam 0.3 mm.

Distribution. Lebanon (type locality: Beirut).

Remarks. No type material was available but the original description was sufficient to identify the species.

Figures

FIGURES 1 – 6. General appearance. 1 — Drilus adustus; 2 — D. akbesianus; 3 — D. bicolor; 4 — D. nemethi sp. nov.; 5 — D. rectus; 6 — D. rufipes. Scale 2.0 mm.

FIGURES 7 – 18. Antenna. 7 — Drilus adustus; 8 — D. akbesianus; 9 — D. bicolor; 10 — D. nemethi sp. nov.; 11 — D. rectus; 12 — D. rufipes. Pronotum. 13 — Drilus adustus; 14 — D. akbesianus; 15 — D. bicolor; 16 — D. nemethi sp. nov.; 17 — D. rectus; 18 — D. rufipes. Scale 1.0 mm (Figs 7 – 12), 0.5 mm (Figs 13 – 18).

FIGURES 19 – 24. Male genitalia. 19 — Drilus adustus; 20 — D. akbesianus; 21 — D. bicolor; 22 — D. nemethi sp. nov.; 23 — D. rectus; 24 — D. rufipes. Scale 0.25 mm.