Gravatamberus pilosus, Mendes & Andersen, 2008

Mendes, Humberto Fonseca & Andersen, Trond, 2008, A review of Antillocladius Saether and Litocladius Mendes, Andersen et Saether, with the description of two new Neotropical genera (Diptera, Chironomidae, Orthocladiinae), Zootaxa 1887 (1), pp. 1-75 : 54-55

Gravatamberus pilosus sp. n.

( Figs 74–78 )

Type material: Holotype male, CHILE: VII Region : Talca, El Golfe, 24–27.xii.1995, Malaise trap, T. Andersen ( ZMBN) . Paratypes: 3 males, same date as holotype; 7 males, same date as holotype except for 27.xx.1995– 2.i.1996; 1 male, Pichilemu Centro, 34º23.121’S, 71º59.983’W, 1 m a.s.l., 17.xi.1998, sweep net, T. Andersen ( MZUSP, ZMBN) .

Diagnostic characters: The species can be separated from all other members of the genus by having more than 10 setae in cell m proximal to RM, costal extension ending close to wing tip, and AR 0.74–0.84.

Etymology: From Latin pilosus , hairy, referring to the strongly hairy wings.

Male (n = 8–10, except when otherwise stated). Total length 1.30–1.50, 1.39 mm. Wing length 0.90–1.12, 1.02 mm. Total length / wing length 1.28–1.47, 1.38. Wing length / length of profemur 2.16–2.39, 2.29 (9). Coloration brown, thorax dark brown without markings, legs uniformly light brown.

Head. AR 0.74–0.84, 0.80. Ultimate flagellomere 220–274, 251 µm long; stout subapical seta 40–65, 51 µm long. Temporal setae 9–12, 10; including 4–6, 5 inner verticals; 2–4, 3 outer verticals; and 2–4, 3 postorbitals. Clypeus with 6–16, 12 setae. Tentorium, stipes, and cibarial pump as in Figure 74 . Tentorium 75–91, 83 µm long; 14–20, 16 µm wide. Stipes 84–98, 91 µm long; 27–34, 30 µm wide. Palp segment lengths (in µm): 14–18, 16; 23–29, 26; 57–79, 68; 66–82, 73; 84–148, 113. Third palpomere with 1–2, 2 sensilla clavata subapically; longest 9–16, 14 µm long.

Thorax ( Fig. 75 ). Antepronotum with 2–3, 3 setae. Dorsocentrals 16–26, 20; acrostichals 9–17, 11, all scalpellate; prealars 6–8, 7, extended anteriorly; supraalar 0–1, 0. Scutellum with 6–9, 7 setae.

Wing ( Fig. 76 ). VR 1.35–1.56, 1.40. C extension 145–168, 157 µm long. Brachiolum with 1 seta; Sc with 19–31, 23 seta; C extension with 25–44, 34 non-marginal setae; R with 18–27, 21 setae; R 1 with 11–19, 14 setae; R 4+5 with 17–29, 23 setae; RM with 0–2, 1 seta; M with 0–2, 1 seta; M with 40–70, 52 setae; M 3+4 with 23–36, 29 setae; Cu with 17–23, 20 setae; Cu 1 with 10–18, 14 setae; PCu with 31–61, 46 setae; An with 21– 28, 25 setae. Cell m with 17–36, 28 setae proximal to RM; r with 350–400 setae; m 1+2 with 400–450 setae; m 3+4 with 10–167, 133 setae; cu + an with 168–262, 210 setae.

1+2

4 + 5

Legs. Spur of foretibia 18–29, 23 (6) µm long; spurs of midtibia 18–27, 22 µm and 11–16, 14 µm long; spurs of hind tibia 32–41, 36 µm and 11–18, 15 µm long. Width at apex of foretibia 20–25, 23 (6) µm; of midtibia 23–25, 24 µm; of hind tibia 25–32, 28 µm. Comb with 8–10, 9 setae; longest 27–38, 33 µm; shortest 16–23, 18 µm long. Lengths and proportions of legs as in Table 12.

Hypopygium ( Figs 77–78 ). Tergite IX with 4–10, 6 setae; laterosternite IX with 3–5, 4 setae. Phallapodeme 48–59, 53 µm long; transverse sternapodeme 52–61, 56 µm long. Gonocoxite 104–118, 111 µm long.

Gonostylus 50–66, 60 µm long; megaseta 4–7, 6 µm long. HR 1.70–2.36, 1.87; HV 2.12–2.88, 2.36.

Biology and distribution: This species is recorded only from Central Chile. The type locality is a boggy area densely grown with 3–4 m high shrubs. The locality at Pichilemu is a river estuary .

Figures

FIGURES 74-78. Gravatamberus pilosus sp. n., male. 74—tentorium, stipes, and cibarial pump; 75—thorax; 76— wing; 77—anal point and tergite IX and dorsal aspect of left gonocoxite and gonostylus; 78—hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right.