HYOTHERIINAE Cope, 1888

Orliac, Maëva J., Karadenizli, Levent, Antoine, Pierre-Olivier & Sen, Sevket, 2015, Small hyotheriine suids (Mammalia, Artiodactyla) from the late early Miocene of Turkey and a short overview of early Miocene small suoids in the Old World, Palaeontologia Electronica 16 (1), pp. 1-18 : 8

Subfamily HYOTHERIINAE Cope, 1888 gen. et species indet.

Description of the Fourth Deciduous Lower Premolar

In addition to n° SMT-1, the Şemsettin locality has yielded a fourth deciduous lower premolar of a suoid (n° SMT-2). The dp4 is typically three-lobed with two cuspids on each lobe. The tooth bears three root pillars: (i) two major ones, an anterior lying under the anterior lobe and the anterior half of the second lobe, and a more massive posterior one under the posterior lobe, and (ii) a smaller labial one, intercalated between the two major root pillars at the level of the protoconid. The tooth only slightly widens posteriorly, and it presents a slight swelling at the level of the protoconid above the small lingual root. The cusps are wide and rounded, separated by narrow and shallow transverse valleys. There is a clear preparaconulid lying between the primonid (sensu van der Made, 1996b) and the paraconid, and connected to them by cristids. The paraconid bears a clear preparacristid, and a postparacristid divided at its base. The primonid has a complex pattern of cristids with pre-, endo-, and postcristids. Postpara- and postprimocristids join the preproto- and premetacristids, respectively, in continuous crests. The cusps of the second lobe (protoconid and metaconid) are heavily worn, but a complicated pattern of crests is still visible, with pre-, post-, and postectocristids. The protoconid also presents an ectoprotocristid. They are linked by their postcristids posteriorly, at the level of the posterior transverse valley. The grooves of the second lobe (pre-, post-,metafossid, and ecto-, pre-, postprotofossid) are deep. The metaconid bears a sharp and high postectocristid that ends in the transverse valley, just in front of the ectoendocristid. Labially, the postectoprotocristid connects to a very light ectohypocristid. The mesoconulid is not clearly differentiated, and it consists in a crest, low in the posterior valley and connected to the prehypocristid. The hypoconid also bears an endohypocristid connected to the endoentocristid and a posthypocristid that joins the small hypoconulid. There is a slight labial cingulum, between the protoconid and the hypoconid.