Glessula mandalayensis Man & Panha, 2024
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publication ID |
https://doi.org/ 10.3897/zookeys.1208.116083 |
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publication LSID |
lsid:zoobank.org:pub:0215503D-E8B4-4179-89F6-FB2CCC59F37E |
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DOI |
https://doi.org/10.5281/zenodo.13137118 |
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persistent identifier |
https://treatment.plazi.org/id/01347449-D63E-469B-826C-BEE5D0D56B8E |
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taxon LSID |
lsid:zoobank.org:act:01347449-D63E-469B-826C-BEE5D0D56B8E |
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treatment provided by |
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scientific name |
Glessula mandalayensis Man & Panha |
| status |
sp. nov. |
14 Glessula mandalayensis Man & Panha sp. nov.
Figs 8 E, F View Figure 8 , 9 View Figure 9 , Tables 1 View Table 1 , 3 View Table 3
Type specimens.
Holotype CUMZ 13072 View Materials (height 14.6 mm, width 7.1 mm; Fig. 8 E View Figure 8 ) , paratypes CUMZ 13073 View Materials (29 shells; Fig. 8 F View Figure 8 ) , NHMUK 20230922 About NHMUK (2 shells) and SMF (2 shells).
Type locality.
Limestone Hills (Apache Cement Factory), Pyinyaung Village, Meiktila District, Mandalay Region, Myanmar (20 ° 49 ' 39.1 " N, 96 ° 23 ' 35.1 " E).
Etymology.
The specific name mandalayensis is the name of region, where the type specimens of this species were collected.
Diagnosis.
Shell large, globosely ovate; spire broad, almost cylindrically attenuated, turreted; last whorl largest; shell surface glossy, smooth to shallow grooves; aperture ovately rounded and broad; peristome thin and white.
Description.
Shell globosely ovate, solid, glossy, brown to bright ochraceous colour, and with 6–7 whorls. Apex blunt and large; protoconch ~ 2 whorls with smooth surface; subsequent whorls with smooth to fine radial striations or sometimes with shallow grooves. Spire broad, cylindrically attenuated, turreted; whorls flatly convex; suture deep; last whorl largest. Aperture ovately rounded and broad; peristome white and thin; columella short, concave, and truncated. Umbilicus closed.
Genitalia (n = 1). Atrium very short. Penis evenly broad, stout, and thick muscularly. Epiphallus short, stout, slightly curved triangular shape, thick muscularly, smooth surface, and ~ 1 / 4 of penis length; flagellum absent. Penial retractor muscle long, thickened and attaches laterally at junction of penis and epiphallus. Vas deferens long distinct tube connected between tip of epiphallus and free oviduct and held in position to penis and vagina with weak connective tissue (Fig. 9 A View Figure 9 ).
Vagina muscularly and nearly as long as penis. Gametolytic duct short and enlarged at base; gametolytic sac small and elliptical in shape. Free oviduct short and stout, and spermoviduct enlarged.
Radula. Each row contains ~ 70 + teeth with half-row formula: central-lateral-marginal teeth (1 – (19–20) – (15–16 + )). Central tooth relatively small, symmetrical tricuspid with pointed central cusp and very small lateral cusps. Lateral teeth bicuspid: endocone long, slender and with pointed tip; ectocone small, pointed tip and located at base of teeth. Latero-marginal teeth gradually reduced in size and asymmetrically bicuspid: endocone short with pointed tip; ectocone small triangle on shape and situated around middle of tooth height. Outermost teeth small and becoming tricuspid; mesocone short and blunt tip; endocone and ectocone very small and pointed tips (Fig. 9 B View Figure 9 ).
Distribution.
This new species is known only from the type locality.
Differential diagnosis.
Glessula mandalayensis sp. nov. can be distinguished from other congeners in Myanmar by having the largest and most globosely ovate shell, obtuse and broad spire, wide aperture, and thinner peristome. In particular, this new species differs from a sympatric species, G. feddeni , by having a larger size (height 13.3–16.0 mm; Table 1 View Table 1 ), more rounded shell, obtuse and broader turreted spire, wider aperture, thinner peristome, and smoother shell surface. In comparison, G. feddeni is smaller in size (height 7.8–10.8 mm; Table 1 View Table 1 ), with a more solid, slender conical shell, pointed and narrow spire, smaller aperture, thicker peristome, and with distinct grooves on the shell surface. Glessula mandalayensis sp. nov. can also be separated from G. latestriata by its broader shell, wider spire, and blunt apex with smoother shell surface. Whereas G. latestriata has a slenderer shell, narrowly attenuated spire and pointed apex with distinct radial grooves striations. This new species also differs from G. gemma by having a larger and globosely ovate shell (height 13.3–16.0; Table 1 View Table 1 ), more turreted spire, convex whorls, vertically elongated aperture, and deeper suture. Whereas G. gemma possesses a smaller and slender shell (height 8.7–9.2; Table 1 View Table 1 ), spire high conical, flatly convex whorls, more rounded aperture, and shallow suture.
Additionally, this new species differs from G. crassilabris s. s. from India and its variety from Rakhine State, Myanmar, in having a smoother shell, obtuse, and lower and broader spire. While G. crassilabris s. s. (Fig. 5 E, F View Figure 5 ) displays stronger striations and a narrowly pointed spire ( Blanford 1865).
Remark.
So far, the genitalia of eight Glessula species has. The terminal part of the male genitalia of G. mandalayensis sp. nov. (Fig. 9 A View Figure 9 ) clearly differs from these eight species in having no flagellum, while a flagellum is well-developed with a comb-like shape in the others (Table 3 View Table 3 ). However, we examined only a single specimen; therefore, the extent of variability within the species remains unknown.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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