Janalia, Janák, 2018

Janák, Jiří, 2018, A new genus of the Tympanophorus lineage of the subtribe Anisolinina from Madagascar (Coleoptera: Staphylinidae: Staphylininae: Staphylinini), Zootaxa 4461 (2), pp. 293-299 : 294

Janalia gen. nov.

Type species. Janalia jarmilae sp. nov., here designated.

Diagnosis. Species of Janalia gen. nov. ( Fig. 1 ) can be recognized within Anisolinina by the following combination of characters: labrum expanded, each lobe less than two times wider then long and not widely separated medially ( Fig. 5 ); apical maxillary palpomere distinctly longer than previous segment, weakly dilated fusiform ( Fig. 6 ), about as broad as previous segment at widest point; apical labial palpomere distinctly dilated, not flattened laterally and with broad apical face ( Figs. 4–6 ); spines on metatibia thin ( Fig. 1 ). Accessory basal lines feebly developed and situated laterally. Aedeagus short and compact, with a short, broad paramere bearing many peg setae ( Figs. 11–14 ).

Description. Form elongate, of medium size. Head small with several macropunctures. Labrum expanded, each lobe less than two times wider then long and not widely separated medially ( Fig. 5 ); Maxillary palpi with apical palpomere distinctly longer than previous triangular segment, weakly dilated fusiform ( Fig. 6 ). Labial palpi with apical palpomere distinctly dilated, not flattened laterally and with broad apical face ( Figs. 4–6 ). Ventral surface of head punctate, mentum markedly transverse ( Fig. 3 ), gular sutures and gular setae distant ( Figs. 3, 4 ). Mandibles relatively short and broad, sharp apically ( Figs. 5, 6 ).

Pronotum with dorsal and lateral series consisting of fine punctures. Prosternum markedly transverse, elevated in middle ( Fig. 3 ), mesoventrital process acute, extending less than half the length of coxae, posteriomedially with slightly protruding ridges ( Figs. 7, 8 ), metaventrite slightly produced between coxae. Elytra with punctuation not forming rows or series. Abdomen with accessory basal lines feebly developed and situated laterally. Legs covered with setae, tibiae with lateral spines.

Male. Sternite 7 in apical third with depression with modified erect setae ( Figs. 9, 10 ). Sternite 8 ( Fig. 9 ) with posterior margin emarginate. Aedeagus short and compact, with broad median lobe bearing many peg setae ( Figs. 11–14 ).

Remarks. The general appearance of the new genus is similar to that of Barygnathus or Trigonopalpus . Janalia gen. nov. shares with Barygnathus the expanded labrum, the aedeagus shape and the distant gular setae, but differs from the latter by the apical labial palpomere not laterally flattened and with a broad apical face. The new genus shares with Trigonopalpus the apical labial palpomere not laterally flattened, with broad apical face, but differs by the latter by the different aedeagus shape and the distant gular setae. In the key to genera of the Tympanophorus lineage, Janalia gen. nov. keys out near Bombylodes and Pammegus (see below), but differs at least from Pammegus (the male of Bombylode s is unknown yet) by short and compact aedeagus with a short, broad paramere. The new genus differs from Bombylodes mainly by different setation patterns on head and by different shape of head, which is strongly trapezoidal in Bombylodes .

Etymology. The genus is dedicated to my wife Jana for her generous long-term support of my activities in entomology. The name is a feminine, consisting of the name Jana and “lia” as a combination of letters.

Figures

FIGURES 1–6. Janalia jarmilae gen. nov., sp. nov. 1, habitus; 2, forebody; 3, head ventral; 4, head ventro-lateral; 5, mouthparts dorsal; 6, mouthparts ventral.

FIGURES 7–14. Janalia jarmilae gen. nov., sp. nov. 7, 8, mesoventrite; 9, sternites 6-8; 10, abdominal segments 6-8 lateral; 11, aedeagus lateral; 12, aedeagus ventro-lateral; 13, aedeagus ventral; 14, aedeagus ventral in transmitted light. 11–14, same scale bars.