Epigambriinae Handlirsch, 1939

Makarkin, Vladimir N., Ansorge, Jörg & Khramov, Alexander V., 2021, Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s. l. (Neuroptera), Palaeoentomology 4 (6), pp. 516-531 : 517

Subfamily Epigambriinae Handlirsch, 1939 , stat. nov.

Epigambridae [sic] Handlirsch, 1913: 813 (nomen nudum); Zittel, 1921: 679 (nomen nudum); Zittel, 1924: 700 (nomen nudum); Labandeira, 2002: 232 (fig. 4).

Epigambriidae Handlirsch, 1939: 76 ; Bode 1953: 18; Brues et al., 1954: 782, 817; Carpenter, 1992: 355; Makarkin & Archibald, 2003: 176; Engel & Grimaldi, 2008: 8 (Table 3); Jepson et al., 2012: 40; Yang et al., 2012: 2.

Type genus. Epigambria Handlirsch, 1939 .

Diagnosis. May be distinguished from members of the polystoechotid genus-group of Ithonidae s.l. by a combination of the following character states: (1) a single gradate series between branches of RP in both fore- and hind wings [two series in genera of the polystoechotid genus-group (rarely: outer series distinct, proximal crossveins numerous, not arranged in inner series in forewing)]; (2) absence of crossveins between branches of MP and CuA in forewing [present in genera of the polystoechotid genus-group]; (3) several crossveins between RA and RP nearly perpendicular to these veins in hind wing [few and oblique in most members of the polystoechotid genus-group].

Included genera. Epigambria , Polyosmylus , and Rasnitsyneura , gen. nov., all from the Early Jurassic (early Toarcian) of Germany.

Remarks. Handlirsch (1913) and Zittel (1921, 1924) mentioned the name ‘Epigambridae’, but no genera were included to this family until 1939.