Rasnitsyneura sp.

Makarkin, Vladimir N., Ansorge, Jörg & Khramov, Alexander V., 2021, Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s. l. (Neuroptera), Palaeoentomology 4 (6), pp. 516-531 : 522-524

Rasnitsyneura sp.

( Figs 6 , 7 )

Material. Specimens PIN 5552/36, PIN 5552/37, PIN 5552/38, PIN 5552/39, PIN 5552/40; incomplete crumpled forewings; all deposited in PIN.

Locality and horizon. Germany: Schandelah. Early Jurassic (early Toarcian, boreale nodule, Harpoceras falciferum ammonite zone).

Description. Specimen PIN 5552/36 ( Fig. 6A, B ). Forewing 5.7 mmlong as preserved (estimated complete length ca. 7–7.5 mm), 2.1 mmwideaspreserved (estimated complete width ca. 2.5). Costal space broadened proximally, distally. Proximal subcostal veinlets bent toward wing apex, simple or with very shallow terminal fork; distal veinlets of Sc, RA mainly once forked. Subcostal space moderately broad distally; crossveins not detected. Sc, RA stout, distally approaching, probably not fused. RA space narrow; crossveins not detected. RP with eight branches, mostly once forked. Three crossveins detected in radial space. MA probably twice forked distally. MP pectinately forked, with three long branches, one of which is deeply forked. CuA pectinately forked, with two very long branches; each branch once or twice shallowly forked. CuP shallowly once forked. A1 pectinately branched, probably with three branches. A2 fragmentarily preserved. Maculation: six brown spots medially (different in size), one paler spot apically.

Specimen PIN 5552/39 ( Fig. 6C, D ). Forewing ca. 7 mmlong as preserved (estimated completelength ca. 7.5 mm), ca. 1.5 mm wide as preserved. Costal space broadened distally. Distal subcostal veinlets of Sc, RA mainly once forked. Subcostal space moderately broad. Sc, RA stout, distally approaching, probably not fused. RA space narrow, with one detected crossveins in middle part. RP with five preserved branches, which mainly once forked distally. CuP once shallowly forked. One crossvein between CuA, CuP detected. A1 pectinately branched, with two/three branches. Maculation: three brown spots preserved medially (all rounded, similar in size), one paler spot apically.

Specimen PIN 5552/38 ( Fig. 6E, F ). Forewing 7.4 mm long as preserved (estimated complete length ca. 8 mm), 2.7 mmwideaspreserved (estimated complete width ca. 3 mm). Costal space relatively narrow, broadened distally. Terminal parts of middle subcostal veinlets strongly bent toward wing apex, with shallow terminal fork; distal veinlets of Sc, RA mostly forked once. Subcostal space narrow as preserved. Sc, RA stout, distally slightly approaching, not fused. RA space narrow; five, irregularly spaced crossveins detected.Anterior trace of RP forked distally, not zigzagged, with probably ten branches. RP1-RP4 dichotomously forked distad outer gradate series; other preserved branches once shallowly forked. One crossvein in radial space detected. Fork of Mnot preserved. MA dichotomously forked distally (like RP1). MP pectinately forked, with five/six long branches (forked once if preserved). CuA pectinately forked, with two, very long branches. CuP shallowly forked once. One (proximal) crossvein between CuA, CuP detected. A1 pectinately forked (branches incompletely preserved). Maculation: five brown spots medially (different in size), one small spot basally, one paler spot apically.

Specimen PIN 5552/40 ( Fig. 6G, H ). Forewing 6.7 mm long as preserved (estimated complete length ca. 7.5 mm), 1.8 mmwideaspreserved (estimated completewidth ca. 2.5 mm). Subcostal veinlets incompletely preserved, bent toward wing apex. RP1, RP2, MA dichotomously forked distad outer gradate series. MP pectinately forked, apparently with four long branches. CuA pectinately forked, with two, very long branches (one branch deeply forked). CuP incompletely preserved, apparently forked once. A1 probably pectinately branched.A2 fragmentarily preserved. Maculation: five brown spots medially (different in size) preserved.

Specimen PIN 5552/37 ( Fig. 7 ). Forewing 6.1 mm long as preserved (estimated complete length ca. 7.6–7.8 mm), 2.4 mmwide aspreserved (estimated completewidth ca. 2.7 mm). Costal space preserved only distally. Distal veinlets of Sc, RA forked once. Subcostal space narrow as preserved. Sc, RA stout, distally approaching, probably not fused. RA space narrow; five/six rather regularly spaced crossveins detected. RP with ten branches. RP1, RP3–RP5, RP10 dichotomously forked distad outer gradate series; RP8 dichotomously forked proximad outer gradate series; RP2, RP9 once shallowly forked. MA rather deeply, dichotomously forked. MP pectinately forked, with four long branches (once shallowly forked). CuA incompletely preserved, pectinately forked, with three/four long branches. Maculation: five dark-brown spots medially (different in size).

Remarks. Thespecimens PIN 5552/36, PIN 5552/38, PIN 5552/39, and PIN 5552/40 may be conspecific and assigned to Rasnitsyneura aequabilis as their forewing maculation, size, and preserved venation are similar to those of specimen 5552/34 (however, the number of the RP branches varies, from seven to ten).

Specimen PIN 5552/37 differs from the others by the absence of apical paler spots and by its broader forewing, suggesting that it may not belong to R. aequabilis .

Figures

FIGURE 6. Rasnitsyneura sp., forewings. A and B, PIN 5552/36. C and D, PIN 5552/39. E and F, PIN 5552/38. G and H, PIN 5552/40. Scale bars = 1 mm (A and B, C–H to same scale).

FIGURE 7. Rasnitsyneura sp., specimen PIN 5552/37. A, Photograph. B, Forewing venation. Scale bar = 1 mm (both to same scale).