Rhyacophila siposi, Kiss, 2013

Kiss, O., 2013, Three New Species Of Rhyacophila (Trichoptera, Rhyacophilidae) From Asia, Acta Zoologica Academiae Scientiarum Hungaricae 59 (1), pp. 13-29 : 19-22

Rhyacophila siposi sp. n.

( Figs 11–15 , 19–20 )

Male: Body length 13.0 mm, forewing length 14.2 mm, forewing width 5.5 mm, length of the antennae 9.0 mm. Body, antennae, palpi, and wings yellowish brown, abdomen brown, legs yellowish brown, genitalia brown ( Figs 19–20 ).

Male genitalia ( Figs. 11–15 , 19–20 ): Apicodorsal lobe of segment IX (a.l.IX, Figs 11–12 ) elongated with downward bent pointed apex in lateral view and obtuse apex with two tiny teeth on either side in dorsal view. Segment X (X, Figs 11–12 ) subtrapezoidal with convex dorsal margin and slightly concave distal margin in lateral view. Anal sclerite (a.s., Figs 11, 14 ) subtriangular with slim-handled lobe. Apical band (a.b., Fig. 11 ) chalice-shaped in lateral view. Proximal part of phallicata (ph., Figs 11, 13 ) thick, thinner in middle, distal part gently curwed downward. Phallicata shorter than first segment of the inferior appendages in lateral view. Paramere (par., Figs 11, 13 ) with three bigger teeth and one tiny tooth dorsally, apex pointed. First segment of the inferior appendages (f.s., Fig. 11 ) long and covered with long bristles. Dorsal lobe of the second joint of inferior appendages (s.j. Figs 11, 15 ) shorter than ventral one, semicircular excision between the dorsal and ventral lobes deep and broad with thick row of tiny teeth in lateral view.

Female: Unknown.

Material – Holotype: male, 3 km SW of Mt. Kalinchok peak, Central Nepal, 27°23’N 86°01’E, 2900 m, by light trapping, 30 June, 1997, leg. Márton Hreblay & Krisztina Csák (gen. prep. No. 117, Ottó Kiss, is deposited in the Mátra Museum, Gyöngyös, Hungary).

Paratype: 1 male, Mt. Kalinchok , Central Nepal, 5 km W of Bigu, 2300 m, 27°63’N 86°09’E, by light trapping, 03 July, 1997, leg. Márton Hreblay & Krisztina Csák, (coll. Ottó Kiss) . 1 male, 6 km SW of Mt. Kalinchok peak, Central Nepal, 3160 m, 27°23’N 86°E, by light trapping, 04 May, 1996, leg. Chenga Sherpa (coll. Ottó Kiss) . 1 male, Mt. Kalinchok , Central Nepal, 8 km E of Barabise, 1850 m, by light trapping 05 July, 1997, leg. Márton Hreblay & Krisztina Csák (coll. Ottó Kiss) .

Diagnosis: This species is similar to Rhyacophila ngorpa Schmid, 1970 ( 1970, p. 133, pl. LIV, figs 16–18; herein Figs 16–18 ), of his R. naviculata group that includes 28 species (about 3.7% of Rhyacophilidae ) described from Asia by Schmid (20 species, 1970), MalicKy (2 species, 1978), Sun and Yang (3 species, 1975), and Kiss (3 species, 2003, 2011).

The characters that place the new species in the R. naviculata group are the second joint of paired inferior appendages, segment X, anal sclerites, apical band, tergal band, phallicata, parameres, and the dorsal appendage.

The most useful characters in helping to recognize the new species are the second joint of paired inferior appendages, apicodorsal lobe of segment IX, anal sclerites, apical band, phallicata, and parameres.

The characters that indicate the relationship of R. siposi sp. n. to R. ngorpa are: − segment X,

− dorsal appendage,

− and tergal band.

The distinguishing characters of R. siposi sp. n. in comparison to R. ngorpa are:

− semicircular excision of the second joint of inferior appendages deeper and broader, upper lobe with obtuse (not acute) distal end;

− apicodorsal lobe of segment IX with a tiny tooth on either side (not without tiny teeth);

− anal sclerite subtriangular with lobe (not quadrangular without lobe);

− apical band chalice-shaped (not horizontally ovate);

− distal part of phallicata slightly curved downward (not horizontal);

− phallicata shorter than first segment of inferior appendages (not as long as first segment of inferior appendages);

− paramere with 4 teeth (not with three ones).

Derivatio nominis: This species is named in memory of my grandmother, Ilona Sipos.

Figures

Figs 11–15. Rhyacophila siposi sp. n., male genitalia: 11 = left lateral view; 12 = apicodorsal lobe of segment IX, dorsal view; 13 = phallic apparatus, left lateral view; 14 = anal sclerite, caudal view; 15 = first segment of inferior appendages and second joint of inferior appendages, left lateral view. (Abbreviations: a.l.IX = apicodorsal lobe of segment IX; a.s. = anal sclerite; a.b. (U) = apical band; C = caudal view; D = dorsal view; d.app. = dorsal appendage; end. = endotheca; f.s. = first segment of paired inferior appendages; L = lateral view; par. = paired paramere; ph. = phallicata; phal. = phallotheca; s.j. = second joint of paired inferior appendages; t. b. (bt) = tergal band; X. = segment X).

Figs 19–20. Rhyacophila siposi sp. n., holotype, male, left lateral view: 19 = habitus, left lateral view, 20 = genitalia, left lateral view.

Figs 16–18. Rhyacophila ngorpa Schmid, 1970, male genitalia (drawings by Schmid 1970): 16 = left lateral view; 17 = apicodorsal lobe of segment IX, dorsal view; 18 = anal sclerite, apical band and segment X, caudal view. (Abbreviations: a.l.IX = apicodorsal lobe of segment IX; a.s. = anal sclerite; a.b. (U) = apical band; C = caudal view; D = dorsal view; d.app. = dorsal appendage; end. = endotheca; f.s. = first segment of paired inferior appendages; L = lateral view; par. = paired paramere; ph. = phallicata; phal. = phallotheca; s.j. = second joint of paired inferior appendages; t. b. (bt) = tergal band; X. = segment X).