Hylodesmum podocarpum (Candolle, 1825) H. Ohashi & R. R. Mill, 2000

Li, Huai-Cheng, Zhao, Xue-Li, Gao, Xin-Fen & Xu, Bo, 2019, Molecular phylogeny of the genus Hylodesmum (Fabaceae), Phytotaxa 403 (3), pp. 221-229 : 227

H. podocarpum group

Normally, this group includes only one species, H. podocarpum ( Candolle 1825: 102) H.Ohashi & R.R.Mill (2000: 181) , which is the type species of the genus ( Ohashi 1973, Ohashi & Mill 2000, Huang & Ohashi 2010). However, there is much debate of the infra-specific classification of the H. podocarpum . Yang and Huang (1979, 1995) thought there were four varieties under the circumstance of the H. podocarpum , i.e., var. podocarpum, var. fallax ( Schindler 1916: 55) Yen C.Yang & P.H.Huang (1979: 8) , var. oxyphyllum ( Candolle 1825: 102) Yen C.Yang & P.H.Huang (1979: 9) , and var. szechuenense (Craib 1914: 104) Yen C.Yang & P.H.Huang (1979: 10) . Ohashi & Mill (2000) and Huang & Ohashi (2010) considered these four varieties should be treated as four subspecies. Gao (2006) believed that the latter two subspecies should be treated as independent species.

Here, we treated them as four subspecies similar to that of Huang and Ohashi (2010). Our study included 18 samples from 4 subspecies. From the ITS analysis and combined ITS and cpDNA analysis, all samples from the Hylodesmum podocarpum group formed a well-supported clade (LB = 100%, PJ = 100%, and PP = 1), which suggested that this group is monophyletic. Besides, each subspecies were individually grouped into a single clade. H. podocarpum subsp. podocarpum and H. podocarpum subsp. fallax (Schindler 1917: 55) H.Ohashi & R.R.Mill (2000: 182) formed a clade with high support values (ITS tree: LB = 95%, PJ = 97%, and PP = 1; combined ITS and cpDNA tree: LB = 99%, PJ = 96%, and PP = 1), while the relationship with the other two subspecies were not well resolved.