Caustogryllacris podocausta pallidior ( Pictet & Saussure, 1891 )

Ingrisch, Sigfrid, 2018, New taxa and records of Gryllacrididae (Orthoptera, Stenopelmatoidea) from South East Asia and New Guinea with a key to the genera, Zootaxa 4510 (1), pp. 1-278 : 25-26

Caustogryllacris podocausta pallidior ( Pictet & Saussure, 1891)

Figs. 3 F–I

Material examined. Indonesia: Java, Magelang , (7°28'53.57''N, 110°13'14.63''E), 1928, leg. Dr. Kühlwein— 1 male (Museum Wiesbaden) .

Description. Small species. Head: Face oval; forehead nearly smooth with very fine impressed dots; fastigium verticis about twice as wide as scapus; ocelli indistinct. Abdominal tergites two and three each with two rows of stridulatory teeth, first row on second tergite with only few teeth, second row sinuate ( Fig. 3J ).

Wings little surpassing hind knees. Tegmen: Radius with a single posterior branch in about mid-length of tegmen, that vein forks in about apical third into radius sector and media anterior; media posterior absent; cubitus anterior forks between basal third and mid-length into two veins, CuA1 and CuA2; cubitus posterior undivided, free throughout; with 4 anal veins, last two with common stem ( Fig. 3F ).

Legs: Fore coxa with a large spine at fore margin; fore and mid femora unarmed; fore and mid tibiae with four pairs of large ventral spines and one pair of smaller ventral spurs; hind femur with 6 external and 2–4 internal spines on ventral margins; hind tibia with spaced spines on both dorsal margins, ventral margins with one preapical spine each; with 3 apical spurs on both sides.

Coloration. General color light brown (specimen formerly conserved in alcohol); vertex unicolored; disc of pronotum indistinctly darker in middle and with dark hind margin. Face black from below compound eyes and including antennal scrobae to clypeo-frontal suture, genae and mouthparts medium to dark brown, the latter with a narrow light band in middle; fastigium verticis, scapus and flagellum yellowish as vertex ( Fig. 3G ). Tegmen semitransparent white with yellow veins and veinlets; hind wing semi-transparent white with yellow veins and brown cross-veins, the cross-veins adorned by narrow light brown bands.

Male. Eighth abdominal tergite moderately prolonged behind. Ninth abdominal tergite moderately prolonged behind with converging and strongly down-bent lateral margins; furrowed in midline, towards tip with a pair of large, strong, elongate swellings that carry on ventral tip a compressed acute tooth ( Fig. 3H ). Tenth abdominal tergite largely reduced. Subgenital plate wider than long with lateral margins in circa basal two thirds convex, afterwards faintly concave, apical area bilobate with obtuse angular excision; a pair of carinae or steps from middle of base to end of convex lateral margins separating convex latero-basal areas fron flattened medio-apical area; styli rather short and stout, inserted at lateral angles of approaching part of margins ( Fig. 3I ). Phallus membranous.

Measurements (1 male).—body w/wings: 18.5; body w/o wings: 15; pronotum: 3.8; tegmen: 14; hind femur: 9; antenna: male> 65 mm.

Discussion. Apart from the lighter coloration, the specimen at hand is similar to C. p. podocausta . The teeth at the bottom of the apical swellings of the male ninth abdominal tergite are compressed and with acute tip while in the specimen from Gunung Arjuna described above these teeth are rounded and with slightly blunt tip. The subspecies of C. podocausta are in need of a revision, including a severe re-examination of the types.

Figures

FIGURE 3. Genus Caustogryllacris: A–E, C. p. podocausta (Haan, 1842) male (A–C) and female (D–E); F–J, C. p. pallidior (Pictet & Saussure, 1891) male.—Habitus dorsal view (A, F), face (B, G), male abdominal apex with ninth tergite and subgenital plate in apico-ventral (C, H) and ventral view (I), stridulatory pegs on second and third abdominal tergites (front is right, J), female subgenital plate (D), ovipositor (E).