Cephennodes (Cephennodes) setifer, Jałoszyński, 2016

Jałoszyński, Paweł, 2016, The Cephenniini of China. VIII. New species and new records of Cephennodes Reitter of Hunan, Jiangxi, Zhejian and Fujian (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 4079 (4), pp. 415-428 : 421-422

Cephennodes (Cephennodes) setifer sp. n.

( Figs 7 , 18–19 )

Type material. Holotype: CHINA (ZHEJIANG Prov.): ♂, two labels: "China: S Zhejiang, Lishui City / Longguan,Fengyang Shan N.R. / mixed forest nr. Luao Vil., 27º / 55' 00''N, 119º11'53''E, moss, / fern, bamboo,bush, sifted, 11 / 30 m, 4.x.2013, Z.-W. Yin leg." [white, printed]; " CEPHENNODES (s. str.) / setifer m. / det. P. Jałoszyński, 2015 / HOLOTYPUS" [red, printed] (SNUC). Paratype: 1 ♂, same data as holotype (cPJ).

Diagnosis. Antennae distinctly shorter than elytra, antennomeres except I and IX–X elongate; elytra with slightly flattened apex around suture, each elytron with several long erect setae but lacking penicilli; median lobe of aedeagus lacking median emargination of ventral apical margin.

Description. Body of male ( Fig. 7 ) strongly convex, oval, with barely marked constriction between pronotum and elytra, brown with slightly lighter legs and distinctly lighter palps, covered with light brown vestiture. BL 1.33 mm.

Head broadest at moderately large but strongly convex and coarsely faceted eyes, HL 0.18 mm, HW 0.30 mm; vertex weakly and evenly convex and anteriorly confluent with frons; frons flattened. Punctures on vertex and frons fine and unremarkable; setae sparse, short, suberect. Antennae short, AnL 0.63–0.65 mm, antennomeres except I, IX and X elongate, IX about as long as broad, X slightly transverse, XI about as long as IX–X together.

Pronotum semi-oval, strongly convex at middle and weakly flattened near hind corners, broadest near middle but barely discernibly narrowing posteriorly; PL 0.43 mm, PW 0.58 mm. Anterior margin in strictly dorsal view slightly arcuate; lateral margins strongly rounded in anterior half and nearly straight in posterior third; posterior pronotal corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae very narrow and demarcated from disc almost on entire length; lateral antebasal pits shallow but distinct, each located indistinctly closer to lateral than to posterior pronotal margin. Punctures on median part of pronotal disc very small but distinct, moderately deep and with sharp margins, those in median area separated by spaces equal to or slightly shorter than their diameters; punctures toward anterior and posterior pronotal margins reducing in size and depth, anterolateral punctures becoming gradually larger, denser and deeper, so that large area adjacent to each anterior corner is covered with distinctly denser and larger but not coarse punctures separated by narrow wrinkles. Setae on pronotal disc moderately long and dense, suberect, additionally pronotum with four pairs of longer and more erect setae: one submedially near anterior pronotal margin, one near middle of lateral margin, one near posterior corner and one submedially near posterior margin. These longer setae are thin and not strikingly long, some of them are difficult to notice.

Elytra about as convex as pronotum and slightly narrower, oval, broadest between middle and anterior third; EL 0.73 mm, EW 0.63 mm, EI 1.16; subhumeral lines carinate but moderately distinct, as long as 0.45× EL and distinctly divergent caudad; basal elytral fovea on each elytron located slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices slightly modified, with small adsutural flattening. Punctures on elytra shallow and fine, unremarkable; setae similar as those on pronotum but slightly longer and more suberect, additionally each elytron with sparse long and strongly erect setae, especially on sides and in posterior third. Hind wings well-developed.

Metaventrite lacking lateral impressions, convex.

Legs long and slender; all tibiae nearly straight.

Aedeagus ( Figs 18–19 ) latus form, AeL 0.28 mm; median lobe in ventral view elongate, broadest near base and gradually narrowing distally; capsular part of median lobe lacking ventral apical emargination; apical projections moderately long, in lateral view dorsal apical projection not curved, pointing distally; parameres moderately broad, each with two robust apical setae.

Female. Unknown.

Distribution. SE China: Zhejiang Province.

Etymology. The specific epithet setifer refers to the long and erect additional elytral setae.

Remarks. Cephennodes setifer is a relatively unremarkable member of the Cephennodes longipes group. It is the smallest species (BL only 1.33 mm; remaining species 1.4 3– 1.63 mm), with only slightly flattened elytral apex around suture (in other species the flattening is either much deeper or entirely absent), and with short antennae in relation to the body. The median lobe of C. setifer lacks the deep median emargination in the ventral apical margin of the capsular part; this character is shared only with C. apicalis Jałoszyński & Nomura, 2009 , C. penicillatus Jałoszyński, 2007a and C. guizhouanus , all of them clearly different in the body shape, proportions of antennomeres and other body parts and in the shape and arrangement of apical projections of the aedeagus. In C. setifer the dorsal apical projection is moderately slender and in lateral view pointing distally, not curved or bent dorsally; in C. apicalis the dorsal apical projection is exceptionally long, slender and distinctly curved dorsally; in C. penicillatus the apex of dorsal apical projection in lateral view is strongly curved dorsally forming a distinct hook; in C. guizhouanus the dorsal apical projection in lateral view is distinctly recurved. Aedeagal diagnostic characters are clearly visible in the holotype of C. setifer despite an erected condition of the copulatory organ, in which the copulatory piece and associated structures are fully extruded.

Figures

FIGURES 5 – 8. Dorsal habitus of males (5, 7, 8) and elytral apex of male in posterodorsal view (6). Cephennodes (s. str.) hystrix sp. n. (5 – 6); Cephennodes (s. str.) setifer sp. n. (7); Cephennodes (s. str.) yinziweii sp. n. (8).

FIGURES 16 – 25. Aedeagus in ventral (16, 18, 20) and lateral (17, 19, 21) views. Cephennodes (s. str.) hystrix sp. n. (16 – 17); Cephennodes (s. str.) setifer sp. n. (18 – 19); Cephennodes (s. str.) yinziweii sp. n. (20 – 21); Cephennodes (s. str.) datianpingensis sp. n. (22 – 23); Cephennodes (s. str.) bicribratus sp. n. (24 – 25).