Zygophylax laertesi, Peña Cantero, 2020
|
publication ID |
https://doi.org/ 10.11646/zootaxa.4822.3.4 |
|
publication LSID |
lsid:zoobank.org:pub:B87B0063-9D1B-4FD7-A2B4-EB3622FCE68A |
|
DOI |
https://doi.org/10.5281/zenodo.4401689 |
|
persistent identifier |
https://treatment.plazi.org/id/F12154FC-7F72-4255-B8C4-0202B01B6413 |
|
taxon LSID |
lsid:zoobank.org:act:F12154FC-7F72-4255-B8C4-0202B01B6413 |
|
treatment provided by |
Plazi |
|
scientific name |
Zygophylax laertesi |
| status |
sp. nov. |
Zygophylax laertesi sp. nov.
( Figs 1D View FIGURE 1 , 3 View FIGURE 3 F–H, 5)
urn:lsid:zoobank.org:act:F12154FC-7F72-4255-B8C4-0202B01B6413
Material examined. MUSORSTOM 4 Stn CP 216, 22°59.5’S– 167°22.0’E (off Koutoumo Island , New Caledonia region), 490–515 m, 29.09.1985: one stem 40 mm high, with coppinia (Holotype, MNHN-IK-2019-2044) GoogleMaps .
Description. Stem 40 mm high ( Fig. 1D View FIGURE 1 ), stiff, strongly and almost completely polysiphonic. Stem with alternately arranged apophyses giving rise to hydrothecae and hydrocladia. Branching alternate, in one plane, but with hydrocladia grouped in sub-opposite pairs ( Fig. 1D View FIGURE 1 , 3F View FIGURE 3 , 5A View FIGURE 5 ) and with two hydrothecae between successive pairs. A basal pair of hydrocladia (and another higher) much developed ( Fig. 1D View FIGURE 1 ), becoming second-order stems or branches and giving rise in turn to hydrocladia. Hydrocladia polysiphonic for most of their extension, except for the youngest ones. Faint separation between hydrocladial apophyses and hydrocladia ( Fig. 3F View FIGURE 3 ). Angle between stem or branches and hydrocladia 70° ( Fig. 1D View FIGURE 1 ). Hydrocladia roughly straight ( Fig. 1D View FIGURE 1 ).
Hydrothecae alternately arranged in one plane ( Figs 3G View FIGURE 3 , 5C View FIGURE 5 ), resting on short apophyses ( Figs 3G View FIGURE 3 , 5C View FIGURE 5 ). Separation between apophysis and hydrothecal pedicel usually marked only by a strong constriction of perisarc ( Figs 3 View FIGURE 3 F–G, 5A–D, F–G), very sharp on adcauline side (occasionally a node present).
Hydrotheca tubiform ( Figs 3 View FIGURE 3 F–G, 5), curved, sigmoid, tapering at base. Hydrotheca bent outwards first, then inwards ( Figs 3G View FIGURE 3 , 5 View FIGURE 5 C–D). Adcauline wall convex at basal half and concave at distal half; conversely, abcauline wall convex at distal half and slightly concave at basal half. Aperture circular, rim even but flared, with up to five renovations. Diameter of hydrothecae roughly constant at distal half, but hydrotheca slightly swollen at basal half. Hydrothecal wall with faint transverse wrinkles. A marked internal adcauline cusp present ( Figs 3G View FIGURE 3 , 5 View FIGURE 5 C–D, F). Hydrothecal diaphragm marked by a strong, oblique ring of perisarc; diaphragm higher at abcauline side ( Figs 3G View FIGURE 3 , 5D View FIGURE 5 , F–G). An internal vertical keel present on abcauline hydrothecal wall above diaphragm ( Figs 3G View FIGURE 3 , 5 View FIGURE 5 C–D, F–G). A dish-shaped operculum present in some hydrothecae ( Fig. 3D, F View FIGURE 3 ).
Two nematothecae resting on hydrothecal and hydrocladial apophyses, one on each side ( Fig. 3G View FIGURE 3 ). Nematothecae on hydrocladial apophyses beyond first hydrothecal apophysis, which is provided with nematothecae ( Fig. 3F View FIGURE 3 , 5B View FIGURE 5 ). Striking abundance of nematothecae on accessory tubes ( Fig. 5 View FIGURE 5 A–B, H), longer and thinner than those on hydrothecal apophyses. Nematothecae elongate, frequently with extra segments (up to five observed) ( Fig. 3F View FIGURE 3 ).
Coppinia rounded ( Fig. 1D View FIGURE 1 ), with an envelope of forked tubes ( Fig. 1D View FIGURE 1 ) covering independent gonothecae. Gonotheca with inverted pear-shaped outline, with a narrow basal part and a much wider distal one, which form two very short arms with a distal aperture ( Fig. 3H View FIGURE 3 ).
Measurements (in µm). Hydrothecae: length of abcauline wall 430–460, length of adcauline wall 380–390, diameter at aperture 75–80, diameter at diaphragm 50–60, length of pedicel 80–100. Nematothecae on apophyses: height 90–100, diameter at aperture 15, maximum diameter 25–30. Nematothecae on accessory tubes: height 170– 250, diameter at aperture 8–15, maximum diameter 20. Gonothecae: height 500–550, diameter at aperture 80–100, maximum diameter 400–500.
Remarks. Zygophylax laertesi sp. nov. is easily recognizable amongst the known species of the genus by the shape of the hydrotheca, the internal adcauline cusp and the abcauline vertical keel above the diaphragm. Zygophylax laertesi sp. nov. resembles no other described species.
Etymology. The specific name laertesi is a dedication to my son Laertes Peña Sancho.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
|
Kingdom |
|
|
Phylum |
|
|
Class |
|
|
Order |
|
|
Family |
|
|
Genus |