Nothogreniera, A.Craig & Currie & Moulton, 2018

A. Craig, Douglas, Currie, Douglas C. & Moulton, John K., 2018, Nothogreniera new genus, for two species of Australian " Paracnephia " (Diptera: Simuliidae), Zootaxa 4444 (4), pp. 351-380 : 352

Nothogreniera gen. nov. Craig, Currie & Moulton

Type species: Simulium fergusoni Tonnoir 1925

Cnephia fergusoni . Mackerras & Mackerras (1949: 386). Generic reassignment.

Paracnephia fergusoni . Crosskey & Howard (1997: 18). Generic reassignment.

Diagnosis. Adults : DARKly PIgMENTED. ANTENNA WITH NINE FlAgEllOMERES , EXTENDED WEll BEyOND HEAD MARgINS; MANDIBlE OF FEMAlE WITH NUMEROUS DISTINCT TEETH ON BOTH SIDES. WINg RElATIVEly lONg IN RElATION TO WIDTH; SMAll BUT DISTINCT BASAl CEll; BASAl RADIAl CEll ElONgATED (A/B RATIO 1.0:2.6); COSTA WITH HAIR-lIKE SETAE ONly ; RS NOT FORKED AND NO EVIDENCE OF TERMINAl EXPANSION, HAIRS ONly; M1 APPARENTly DOUBlE (PRODUCED By PARAllEl FOlD WITH CONCENTRATION OF MICROTRICHIA) ; CUA MARKEDly SINUOUS. ANEPISTERNAl (PlEURAl) MEMBRANE lACKINg HAIRS; ANTEPRONOTAl lOBE AND PROEPISTERNUM HAIRED; KATEPISTERNAl SUlCUS POORly EXPRESSED. HIND BASITARSUS WITH VENTRAl ROW OF SPINES; CAlCIPAlA ABSENT TO POORly DEVElOPED, AggREgATION OF STOUT SPINES IN THAT POSITION; PEDISUlCUS ABSENT; TARSOMERE II ElONgATE; ClAWS OF FEMAlE WITH TOOTH ABSENT, OR IF PRESENT, MINUTE, ROUNDED HEEl WEll DEVElOPED. HyPOgyNIAl VAlVES WEll STRENgTHENED AlONg MEDIAl EDgES; gENITAl FORK ANTERIOR ARM THIN, DISTINCT, lATERAl ARMS lACKINg ANTERIOR POINTINg APODEMES; SPERMATHECA SPHERICAl, SURFACE SMOOTH, SPARSE INNER ACANTHAE, DUCT JUNCTION SCUlPTED, ClEAR AREA SMAll. MAlE MARKEDly HIRSUTE; VENTRAl PlATE OF gENITAlIA HIRSUTE, BROAD AND FlATTENED APICAlly WITH SlIgHT lIP, MEDIAl CARINA POOR-TO-WEll EXPRESSED, BASAl ARMS SUBSTANTIAl AS ARE PARAMERE CONNECTORS, PARAMERES AS FlATTENED PlATES THICKENED AlONg ONE EDgE , SPINES ABSENT, ADEAgAl MEMBRANE WITH CONCENTRATION OF SMAll SPINES CENTRAlly; gONOSTylI BROAD, TWO SUBSTANTIAl SHORT TERMINAl SPINES. Pupa : gIll WITH 15 OR 16 FINE UNBRANCHED FIlAMENTS ; ABDOMEN WITH ANTERIOR SMAll SPINE COMBS ON TERgITES VI–IX, lIgHTly EXPRESSED ON TERgITE V; MODERATE PlEURITE ON SEgMENT V, SMAllER PlEURITES ON SEgMENTS VI AND VII; SEgMENT IX WITH TWO SHORT SUBSTANTIAl TERMINAl SPINES PlUS gRAPNEl HOOKS. Cocoon : SHAPElESS, FABRIC OF COARSE FIBRES, COVERINg ONly POSTERIOR PORTION OF ABDOMEN, OFTEN WITH EXTRANEOUS MATERIAl . Larva : OVERAll DARK BROWN OR yEllOWISH, lABRAl FANS NORMAl TO ENlARgED ; ANTENNA SUBEQUAl TO, OR lONgER THAN FAN STEM, BASAl AND MEDIAl ANTENNOMERES SUBEQUAl IN lENgTH, TOgETHER SUBEQUAl TO DISTAl ANTENNOMERE; APICAl ANTENNOMERE WITH FINE BANDINg PATTERN OBSERVABlE AT HIgH MAgNIFICATION; CEPHAlIC SETAE NOT EXACERBATED; HyPOSTOMA WITH TEETH IN THREE DISTINCT gROUPS, ONE MAIN HyPOSTOMAl SETA PER SIDE; MANDIBlE WITH ENlARgED PROXIMAl PREAPICAl TOOTH, SERRATION AND SENSIllUM WEll DEVElOPED; gENAl ClEFT MARKEDly SHAllOW V-SHAPED TO DEEPER, POORly EXPRESSED; CERVICAl SClERITES FUSED TO POSTOCCIPUT; lATERAl SClERITE OF ANTERIOR PROlEg D-SHAPED; VENTRAl TUBERClES ABSENT TO POORly EXPRESSED; ANAl SClERITE OF POSTERIOR PROlEg X-SHAPED, NOT MARKEDly EXPRESSED, INTERARM STRUTS ABSENT; RECTAl SCAlES ABSENT; CIRClET OF HOOKS WITH SMAll NUMBER OF ROWS AND HOOKS.

Etymology. FROM LATIN nothus (= FAlSE OR BASTARD).

Constituents. Nothogreniera fergusoni ( TONNOIR, 1925) (TyPE SPECIES), N. occidentalis ( MACKERRAS & MACKERRAS, 1949).