Dactylonotus tsitsikamma, Igor Ya. Grichanov, 2016

Igor Ya. Grichanov, 2016, Two new species of Dactylonotus Parent, 1934 (Diptera: Dolichopodidae) from South Africa and a key to Afrotropical species, European Journal of Taxonomy 175, pp. 1-9 : 4-7

Dactylonotus tsitsikamma sp. nov.

urn:lsid:zoobank.org:act:9D497D9A-DFC2-4856-857A-86C75C373BB6

Figs 5–10, 12

Diagnosis

The new species is close to D. univittatus (Loew, 1858) as described by Grichanov (2000), differing in the presence of the posterior brush of long setae on the fifth segment of the fore tarsus, the brownish yellow hypopygium and the morphology of the hypopygial appendages. The arista-like stylus is middorsal; the fore coxa are with black hairs; the mid femur is without long hairs and setae.

Etymology

The species is named after the Tsitsikamma National Park in South Africa, where the type material was collected. Type material

Holotype

REPUBLIC OF SOUTH AFRICA: ♂, [Eastern Cape,] Tsitsikamma N.P., below Sleepkloof , malaise trap, indigenous forest, 33 ° 56.974 S, 23 ° 54.926 E, 20–22 Jan. 2009, 64 m, A.H. Kirk-Spriggs & S. Otto ( BMSA).

Description

Male (Figs 5–10)

MEASUREMENTS. Body length without antennae 5.5 mm, antenna length 1.6 mm, wing length 5.0 mm, wing width 1.7 mm. HEAD (somewhat shrunken). Frons as wide as high, black, grey pollinose, with medial rhomboid depression; face densely whitish pollinose, narrow; occiput flat, black, grey pollinose; one pair of long ocellar and short postvertical setae; no vertical setae; postocular setae black in upper part and white in middle and lower parts of head; eyes with short white hairs; antennae (Figs 6–7) inserted at upper fourth of head, dirty-yellow, brownish dorsally and apically, 1.5 × longer than height of head; scape long, bare; pedicel long, slightly widened at apex, dorso-lateral (inner view) process, covered by dorsal, lateral and ventral setulae; postpedicel very long, widest at apex of pedicel, with narrow rounded apex, 2.8 × as long as high in middle, entirely covered by microscopic hairs; arista-like stylus middorsal, with short hairs. Length ratio of scape to pedicel to postpedicel (dorsal to ventral sides) to arista-like stylus (1st and 2 nd segments) in mm, 0.33/0.38/0.40/0.73/0.43/0.60. Palpus and proboscis short, yellow, with black hairs.

THORAX. Mostly brownish yellow, with black bristles; mesonotum with broad median metallic blue violet stripe embracing area between 2 nd–6th pairs of dorsocentrals; scutellum dorsally blue-violet with yellow margin; pleura with a black spot below calypter; thoracic pollination weak; proepisternum with 2 long brownish ventral setae and 4 yellow hairs above; 6 pairs of dorsocentral bristles decreasing in size anteriorly, with several scattered hairs on anterior slope; acrostichals biseriate, extending to 5 th pair of dorsocentrals; scutellum with two long strong bristles and two short fine lateral setae, 1/4 as long as medians, dorsally bare.

LEGS. Including coxae yellow, with black bristles, apical segments of all tarsi brownish; fore coxa anteriorly with short black hairs and row of six black lateral and apical setae of various length; mid coxa anteriorly with rather long black hairs and setae; mid and hind coxae with black bristle at base; hind coxa with additional small seta below middle; femora without long setae; mid and hind femora each with one strong preapical anterior bristle; fore tibia with 3 strong anterodorsal, 2 short posterodorsal setae; mid tibia with 3 strong anterodorsal, 1 small dorsal, 3 strong posterodorsal setae; hind tibia with 3 anterodorsal, 5 posterodorsal, 4–5 short ventral setae; all tibiae with strong apical setae; 4 th and 5 th segments of fore tarsus slightly swollen; 5 th segment (Fig. 9) of same tarsus with posterior brush of flattened setae, as long as 5 th segment, with slightly enlarged claws and pulvilli; mid and hind tarsi simple, with short claws and small pulvilli. Femur, tibia and tarsomere (from first to fifth) length ratio (in mm): fore leg: 1.65/1.63/1.02/0.41/0.28/0.2/0.22, mid leg: 1.82/2.05/1.14/0.55/0.43/0.28/0.19, hind leg: 2.27/2.5/0.7/0.71/0.42/0.27/0.19.

WING (Fig. 8). Ovate, almost hyaline; veins brown; R2+3 and R4+5 slightly curved posteriad at apex; M1+2 nearly straight; crossvein dm-cu straight; ratio of dm-cu to apical part of CuA 1 (in mm) 0.36/0.92; anal vein foldlike; anal angle obtuse; lower calypter yellow, with black cilia; halter yellow.

ABDOMEN. Mostly yellow, with black hairs and marginal setae; 1 st segment brownish anteriorly; 2 nd segment with brown T-shaped spot dorsally; 3 rd– 5 th segments each with triangular dark-brown spot dorsally; 6 th segment reduced, brown; 7 th segment small, bare; 8 th segment and epandrium mostly dirty yellow, partly brown, 8 th segment with short hairs and row of strong setae; hypopygium ( Figs 10, 12) yellow-brown; epandrium ovate; hypandrium broad, with 2 large ventral projections; phallus pointed at apex; two epandrial setae positioned on distal side of epandrium, one of them (dorsal) pedunculate; epandrial lobe long, fingerlike, with rounded apex and 2 short apical setae; surstylus yellow-brown, bilobate, with almost straight lobes, with several apical setae as figured; ventral lobe thinner than dorsal lobe, clavate; postgonite thick, slightly curved ventrally; cercus short, yellow, with black setae.

Female

Unknown.

Distribution

South Africa (Eastern Cape).