Mioscarta nubisa Crispolon & Soulier-Perkins, 2021
publication ID |
https://doi.org/10.5852/ejt.2021.778.1571 |
publication LSID |
lsid:zoobank.org:pub:7A844BA1-CBC8-437D-8110-E1C598778057 |
DOI |
https://doi.org/10.5281/zenodo.5705002 |
persistent identifier |
https://treatment.plazi.org/id/17979C2C-D071-44EA-9D16-974A742808A1 |
taxon LSID |
lsid:zoobank.org:act:17979C2C-D071-44EA-9D16-974A742808A1 |
treatment provided by |
Felipe (2021-11-16 13:13:07, last updated 2024-12-04 22:17:16) |
scientific name |
Mioscarta nubisa Crispolon & Soulier-Perkins |
status |
sp. nov. |
Mioscarta nubisa Crispolon & Soulier-Perkins sp. nov.
urn:lsid:zoobank.org:act:17979C2C-D071-44EA-9D16-974A742808A1
Fig. 7 View Fig
Diagnosis
General shape of M. nubisa is similar to M. obscuripennis but are distinctly different in color. M. nubisa in dorsal view presents a brownish and opaque tegmen, darker toward apex, pronotum brownish and yellowish or brown legs while M. obscuripennis has a dark brown or black tegmen, brown or orange pronotum and orange legs.
Etymology
Species name refers to the light to darker coloration forming a cloudy pattern on the tegmen and is the female superlative of the latin word “nubis” which means cloudy.
Material examined
Holotype PHILIPPINES • ♂; “Philippines, Camarines Sur, Luzon isl. Mt Isarog Natural Park , Panicuason Naga ”; “Muséum Paris; 1 May 2011; S. A. Yap, M.V. Yngente, O.L. Eusebio rec.”; UPLBMNH HEM-05052 ”; UPLBMNH.
Paratypes PHILIPPINES • ♀; “ Trinidad ; 22 June 1951; C.R. Baltazar ”; “UPLBMNH HEM 04064”; UPLBMNH • ♂; “ Philippines, Camarines Sur, Luzon isl. Mt Isarog Natural Park , Panicuason Naga ”; “Muséum Paris, 1 May 2011; S.A. Yap, M.V. Yngente, O.L. Eusebio rec.”; UPLBMNH HEM-05053 ”; UPLBMNH • 16 ♂♂; “ Philippines, Luzon, mount Isarog , Light trap ”; “Muséum Paris; 31 May 2011; S. Yap Rec. ”; “Muséum Paris, MNHN ( EH) 23645 to 23646 , 23648 , 24748 , 24859 to 24870 ”; MNHN • ♂; “ Philippines, Luzon isl , Laguna, Los Banos , Mount Makiling ; 14º082' 29" N, 121º12.242" E; “Muséum Paris; 7 Nov. 2008; 605 m, Primary forest, at sight, A. Soulier-Perkins, S. Yap Rec. ”; “Muséum Paris, MNHN ( EH) 23644 ”; MNHN .
Description
BODY. Length 10 mm (tegmina included), width 5 mm.
HEAD ( Figs 1A View Fig , 7B View Fig ). In dorsal view, large ocelli, distance between eyes less than 8 times ocellus diameter, distance between ocelli equals one ocellus diameter, distance between ocellus and compound eye 2 times ocellus diameter, ocelli closer to each other than from compound eyes. Eyes not prominent, length 1.44 times than wide. Vertex and frons longitudinal median carina absent. Vertex slightly longer than wide with 3 times ocellus diameter in between two vertex grooves outside ocelli and 3.5 times ocellus diameter between anterior and posterior vertex margins. Postclypeus with longitudinal furrow, slightly swollen and ovoid shape in frontal view, widest part at mid height ( Figs 1A View Fig , 7B View Fig ), not receding and prior to anteclypeus where it bends forming obtuse angle in lateral view ( Fig. 7A View Fig ). Rostrum long, reaching but not surpassing mesocoxae. Thorax ( Figs 2A View Fig , 7A–C View Fig ). In dorsal view, pronotum with anterior concavities on each side, anterior margin of pronotum as wide as posterior margin of head including eyes, anterolateral margin curved, posterior margin grooved, postero-lateral margin slightly concave, longer than anterolateral margin, humeral angle rounded. In lateral view, pronotum curving not more than 25º ( Figs 2 View Fig , 7A View Fig ). Scutellum as long as wide with large median dimple ( Fig. 7C View Fig ). Tegmen ( Fig. 7A–C View Fig ). R bifurcates on apical half, M bifurcate on basal third, apical reticulation well developed without concave apical cells. Posterior wing ( Fig. 3A View Fig ), Rp separating from SC+Ra nearly at midlength, M reaches ambient vein, Cua and Cup fused at base and m-cu links M to Cua before Cua bifurcation, common base for Cup and Cua originate at base of wing, posterior wing with 7 longitudinal veins and 5 apical cells between SC+Ra and Cup, angular protrusion of costal margin near its base present. Metafemur with apical spine in inner margin, metatibiae bearing 1 lateral spine.
MALE TERMINALIA. In lateral view, posterior margin of pygofer largely convex in middle with slight curved on last third ( Fig. 7E View Fig ). Subgenital plates ( Fig. 7F View Fig ) very long relative to height of pygofer with fine tapering appendage longer than main plate, dorsal and ventral margin of main plate straight, sterno-lateral plate present, slightly elongated. Intermediate plate present, elongated, roughly oblong shaped, linking internal sides of lateral and subgenital plates. Paramere ( Fig. 7G View Fig ) not globose, dorsal margin convex and regularly curving finishing by a sharp process pointing dorsally, ventral margin convex and largely angled subapically, apex with spiniform process pointing postero-ventrally. Aedeagus ( Fig. 7H View Fig ) with basal third of dorsal margin regularly bent without angle before the bent part, last 2 ⁄ 3 vertical and S-shaped, ventral margin regularly curved, apical extension pointing posteriorly, posterior protrusion axe-shaped with edge prolonged ventrally by a straight, long and thin extension, postero-dorsal protrusion absent.
COLOR. Head, antennae, pronotum, abdomen, legs scutellum light brown, rostrum brownish highlighted with white. Tegmen opaque light brown to brown.
Type locality
Philippines: Camarines Sur, Luzon isl. Mt Isarog Natural Park, Panicuason Naga.
Distribution
Philippines: Luzon Island.
Astrin J. J. & Stuben P. E. 2008. Phylogeny in cryptic weevils: molecules, morphology and new genera of western Palaearctic Cryptorhynchinae (Coleoptera: Curculionidae). Invertebrate Systematics 22 (5): 503 - 522. https: // doi. org / 10.1071 / IS 07057
Colgan D. J., McLauchlan A., Wilson G. D. F., Livingston S. P., Edgecombe G. D., Macaranas J., Cassis G. & Gray M. R. 1998. Histone H 3 and U 2 snRNA DNA sequences and arthropod molecular evolution. Australian Journal of Zoology 46: 419 - 437. https: // doi. org / 10.1071 / ZO 98048
Cryan J. R., Wiegmann B. M., Deitz L. L. & Dietrich C. H. 2000. Phylogeny of the treehoppers (Insecta: Hemiptera: Membracidae): evidence from two nuclear genes. Molecular Phylogenetics and Evolution 17: 317 - 334. https: // doi. org / 10.1006 / mpev. 2000.0832
Dietrich C. H., Rakitov R. A., Holmes J. L. & Black W. C. 2001. Phylogeny of the major lineages of Membracoidea (Insecta: Hemiptera: Cicadomorpha) based on 28 S rDNA sequences. Molecular Phylogenetics and Evolution 18: 293 - 305. https: // doi. org / 10.1006 / mpev. 2000.0873
Hillis D. M. & Dixon M. T. 1991. Ribosomal DNA: molecular evolution and phylogenetic inference. The Quarterly Review of Biology 66: 411 - 453. https: // doi. org / 10.1086 / 417338
Schmidt E. 1925. Funf neue Zikadenarten (Forts f.). Societas Entomologica. Organ fur den internationalen Entomologenverein 40: 35 - 36.
Whiting M. F., Carpenter J. C., Wheeler Q. D. & Wheeler W. C. 1997. The Strepsiptera problem: phylogeny of the holometabolous insect orders inferred from 18 S and 28 S ribosomal DNA sequences and morphology. Systematic Biology 46: 1 - 68. https: // doi. org / 10.1093 / sysbio / 46.1.1
Whiting M. F. 2002. Phylogeny of the holometabolous insect orders based on 18 S ribosomal DNA: when bad things happen to good data. In: DeSalle, R., Giribet, G., Wheeler, W. (ed.), Molecular Systematics and Evolution: Theory and Practice, Birkhauser, Basel: 69 - 83. https: // doi. org / 10.1007 / 978 - 3 - 0348 - 8114 - 2 _ 5
Fig. 7. Mioscarta nubisa Crispolon & Soulier-Perkins sp. nov. holotype, habitus and ♂ terminalia in lateral view. A. Lateral view. B. Frontal view. C. Dorsal view. D. Labels. E. Terminalia. F. Sternolateral, intermediate and subgenital plates. G. Paramere. H. Aedeagus.
Fig. 1. Head schematic representation with diameter of ocelli compared to distance between them and between ocellus and compound eye. A. Mioscarta Breddin, 1901. B. Trigonoschema Crispolon & Soulier-Perkins gen. nov. C. Poeciloterpa Stål, 1870.
Fig. 2. Schematic representation of the pronotum angle. A. Mioscarta Breddin, 1901. B. Trigonoschema Crispolon & Soulier-Perkins gen. nov. C. Poeciloterpa Stål, 1870.
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