Typhlopsychrosoma

Gilgado, José D., Enghoff, Henrik & Ortuño, Vicente M., 2015, The hypogean Iberian genus Typhlopsychrosoma Mauriès, 1982 (Diplopoda, Chordeumatida, Vandeleumatidae): distribution map, key to species, first record in a Mesovoid Shallow Substratum (MSS) and detailed iconography of T. baeticaense (Mauriès, 2013), Zootaxa 3937 (2), pp. 337-346 : 343

Key to the species of Typhlopsychrosoma (males)

1 Distal process c of part C of colpocoxite very long and slender, (maximum length from the tip of c to the elbow of C more than 7 times the width), uniformly decreasing in width towards the tip as seen in lateral view. Bundle of bristles (‘lanières’) f large and conspicuous, resembling a fan, reaching the tip of c, and more than 4 times as wide as the width of c at half its length. Syntelopodite similar in size to colpocoxite. Adult body length> 15 mm ............................................ 2

- Distal process c of part C of colpocoxite not so long and slender (maximum length from the tip of c to the elbow of C less than 7 times the width), but rather robust, as seen in lateral view. Bundle of bristles f narrow and dense, reaching the tip of c or not. Syntelopodite inferior in size to colpocoxite, embraced by it. Adult body length <15 mm ........................... 3

2 Elbow of part C of colpocoxite without a conspicuous protrusion, distal part of colpocoxite process c uniformly curved with no conspicuous angle. Lateral parts of syntelopodite t not curved ( Fig. 4 E)................................................................................... T. fadriquei from type locality (Cueva del Oro, Cornión mountains, Asturias)

- Elbow of part C of colpocoxite not protruding, distal part of colpocoxite process c with a curved angle. Lateral parts of syntelopodite t patently curved ( Fig. 4 F)......................................................................... T. fadriquei from Cave “Hoya la Madre”, Ravine of River Casaro, Cornión mountains, Asturias. Possibly a malformed specimen, according to Mauriès (1982)

3 Tip of the process c of part C of colpocoxite curved at an angle of almost 90 degrees, projecting towards the gonopod basis. Tips of lateral parts of syntelopodite t blunt as seen in lateral view ( Fig. 4 D)................................................................................................... T. tarraconense (Cova Cambra, Tortosa, Tarragona)

- Tip of the process c of part C of colpocoxite with no angle, although a distal keel may be present dorsally. Tips of lateral parts of syntelopodite t finishing in point ( Fig. 4 A,C)............................................................ 4

4 Part C reaching basis of colpocoxite, part K of colpocoxite placed mostly anteriorly. Lateral parts of syntelopodite t long, reaching (or almost) the posterior part of the elbow of C, with a soft curvature in an angle slightly superior to 90 degrees as seen in lateral view ( Fig. 4 A,B)...................................................................................... T. baeticaense (several locations in the Baetic System Mountain Range, Córdoba, Jaén, Alicante [new records]).

- Part C not reaching basis of colpocoxite, but resting on part K, which is placed not only anteriorly but also lateromedially. Lateral parts of syntelopodite t short, almost but not reaching half the distance from the base of colpocoxite to the elbow, and with a curvature of almost 180 degrees as seen in lateral view ( Fig. 4 C)......................................................................................................... T. breuili (Atapuerca Caves, Ibeas de Juarros, Burgos).

Figures

FIGURE 4. Gonopods of all Typhlopsychrosoma species in lateral view: A) T. baeticaense from type locality; adapted from Mauriès (2013). B) T. baeticaense from Aitana Mountains. C) T. breuili; adapted from Mauriès (1971). D) T. tarraconense; adapted from Mauriès (1971). E) T. fadriquei from the type locality; adapted from Mauriès & Vicente (1978). F) T. fadriquei from Hoya La Madre cave (possibly an anomalous specimen according to Mauriés (1982); adapted from Mauriès (1982).