Harperocallis falcata (Ruiz & Pav.) L.M. Campb. & Dorr, 2013

Campbell, Lisa M. & Dorr, Laurence J., 2013, A synopsis of Harperocallis (Tofieldiaceae, Alismatales) with ten new combinations, PhytoKeys 21, pp. 37-52 : 39-41

Harperocallis falcata (Ruiz & Pav.) L.M. Campb. & Dorr comb. nov.

Isidrogalvia falcata Ruiz & Pav., Fl. Peruv. 3: 69, t. 302b. 1802. Type: Peru. de Pillao [sic], H. Ruiz & J.A. Pavon s.n. (lectotype: MA [MA810531!], F-negative no. 29435 (F!, MO!), selected by Cruden, 1991; possible isolectotypes: BC [BC872749!], BM [BM000938089!], MA [MA810498!, MA [MA810530!]).

Tofieldia falcata (Ruiz & Pav.) Pers., Syn. Pl. 1: 399. 1805. Type: Based on Isidrogalvia falcata Ruiz & Pav.

Tofieldia falcata (Ruiz & Pav.) Willd., Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 2: 29. 1808, comb. illeg. Type: Based on Isidrogalvia falcata Ruiz & Pav.

Narthecium falcatum (Ruiz & Pav.) Poir., Encycl., Suppl. 4: 61. 1816. Type: Based on Isidrogalvia falcata Ruiz & Pav.

Asphodeleris falcata (Ruiz & Pav.) Kuntze, Revis. Gen. Pl. 2: 706. 1891. Type: Based on Isidrogalvia falcata Ruiz & Pav.

Tofieldia flexuosa Willd., Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 2: 28. 1808. Type: Peru [Ecuador?]. F.W.H.A. von Humboldt & A.J.A. Bonpland 3383 (holotype: B-W [B-W-07103-01 0!]; isotypes: GH-fragm., P [P02137266!]).

Tofieldia frigida Kunth in H.B.K., Nov. Gen. Sp. [quarto ed.] 1: 267. 1815 [1816]; Ibid. [folio ed.] 1: 213. 1815 [1816]. Type: Peru [Ecuador?]. F.W.H.A. von Humboldt & A.J.A. Bonpland 3383 (neotype, here designated: B-W [B-W-07103-01 0!]; isoneotypes: GH-fragm., P [P02137266!]).

Type.

Based on Isidrogalvia falcata Ruiz & Pav.

Distribution

( Fig. 1 ). The Andes of Ecuador (Azuay, Cañar, Loja, Morona-Santiago, and Zamora-Chinchipe provinces) and Peru (Amazonas, Cajamarca, Cusco, Huánuco, Junín, and Pasco regions); 2300-3860 m.

Representative specimens.

ECUADOR. Azuay: Cordillera Oriental, alrededores del Páramo de Patococha entre Gualaceo y Limon, 3400-3450 m, 6-7 Aug 1959, H.G. Barclay & P. Juajibioy 8632 (MO, NY); Km 85 on Pan American Highway N of Loja, 2850-2950 m, 3 May 1973, L. Holm-Nielsen et al. 4815 (MO, NY); Hac. Horta-Naque, 3100 m, 11 Jun 1946, H.N. Moldenke 869 (NY). Cañar: Cerro Yanguán NE of Pindilig, 3100 m, 13 Dec 1980, L.B. Holm-Nielsen et al. 29300 (MO, NY). Loja: Loma de Oro at Panamerican Highway, 3300 m, 2 Jan 1981, H. Balslev 1382 (MO, NY, US); Yangana-Zumba road Km 15-20, N slopes of Cordillera de Sabanilla, 2550 m, 31 Dec 1980, H. Balslev 1298 (NY). Loja/Zamora-Chinchipe: Parque Nacional Podocarpus, crest of the Cordillera de los Andes E and SE of Nudo de Cajanuma, s.d., B. Øllgaard 90768 (NY). Morona/Santiago: Gualaceo-Sigsig-Gualaquiza road, SSE of Sigsig, 03°11'S, 78°40'W, ca. 2900-3090 m, 3 Dec 1990, J.L. Luteyn et al. 14287 (NY). PERU. Amazonas: Prov. Chachapoyas: Cerca a Calla-Calla, siguiendo la ruta a Leymebamba-Balsas, 3860 m, 18 Dec 1992, I. Sanchez Vega & J. Tanta 6481 (US); Cerros [de] Calla Calla, east side, 19 km above Leimebamba [sic] on the road to Balsas, 3100 m, 4 Jun 1964, P.C. Hutchinson & J.K. Wright 5510 (NY, US). Cajamarca: Prov. Cutervo: 2350 m, 22 Jun 1992, I. Sanchez Vega & A. Miranda 6265 (NY).

Notes.

When compared to its present on-line image (http://plants.jstor.org/specimen/ma810531), a photograph (F-negative no. 29435) of the lectotype of Isidrogalvia falcata taken by J. Francis Macbride of the Field Museum before World War II shows that the original label (" Isidrogalvia falcata Sp. Pl. Fl. Per. de Pillao") was moved from the center of the specimen to the lower left-hand corner.

Tofieldia frigida was based on a collection or collections made by F.W.H.A. von Humboldt & A.J.A. Bonpland in Ecuador ("Crescit in frigidis regni Quitensis inter Loxam et pagum Ona, in summis montibus Saraguri, et Alpachacae, alt. 1200-1400 hexap, … Floret Decembri."). While no material with this name was found in Paris (P-Bonpl.) ( Stauffer et al. 2012), the type specimen of Tofieldia flexuosa in B-W is annotated as Isidrogalvis [sic] frigida Klotzsch, nom. nud. This strongly supports that the names Tofieldia frigida and Tofieldia flexuosa are based on the same gathering (i.e., F.W.H.A. von Humboldt & A.J.A. Bonpland 3383) and we have selected this collection to neotypify the name Tofieldia frigida .

Figures

Figure 1. Distribution of Harperocallis.