Sulciclivina curvata, Balkenohl, 2023

Balkenohl, Michael, 2023, Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini), European Journal of Taxonomy 915 (1), pp. 1-57 : 39-42

Sulciclivina curvata sp. nov.

urn:lsid:zoobank.org:act:47AF8CF1-C02B-4537-9826-B91C368A6A86

Figs 15 , 31, 46, 53

Diagnosis

A medium-sized species with a long-oval shape of the elytron, interval eight of the elytron covered by half of the width with indistinct reticulation, and the lateral lobe completely fused with the wing. The most similar species are S. sulcigera and S. striata striata . In S. sulcigera , the reticulation of interval eight is distinct and very often extended to interval seven. In addition, the proepisternum is distinctly visible in dorsal view and interval eight is carinate up to the tip of the apex. In the more slender species S. striata striata , the pronotum is longer than wide and with the margins subparallel, the median part of the clypeus is indistinctly bilobed and nearly straight. Moreover, the aedeagi and the gonocoxites of these species all show unique characters.

Etymology

The name refers to the curved and distorted apical third of the gonocoxite by the Latin adjective for bent in the feminine form ( ʻ curvata ʼ).

Type material

Holotype

INDONESIA • ♂; with small mounting card, with labels and data: white, black printed and handwritten in black ink “Batoerraden G. Slamat. Java F.C.Drescher. 10-12 VII 1926 ” / white, handwritten in black ink “ C. attenuata Hbst Andr. det. 23.11.1933 ” / white, handwritten in black ink and black printed “C. pseudoattenuata m. det. K. Kult 44”; ETHZ.

Remark. The holotype is slightly immature.

Taxonomic remark

Kult recognized the form as being a new species and added a name to the specimens. However, it was never described and the name was never published. The name provided to the specimen by Kult is therefore not valid.

Paratypes

INDONESIA • 1 ♀; “F.C.DRESCHER Djeroeklegi Zuid-Banjoemas 9.XI.1930 ” / “attenuata Herbst Andr. det. 23.11.33 ”; ETHZ • 1 ♂; “F.C.DRESCHER Djeroeklegi Zuid-Banjoemas 9.XI.1930 ” / “ C. sulcigera Putz. ?” / “attenuata Hbst Andr. det. 23.11.33 ” / “COLLECTIO KAREL KULT COLL. A.DOSTAL, 1999”; CADW • 1 ♂; “F.C.DRESCHER Djeroeklegi Zuid-Banjoemas 9.XI.1930 ” / “attenuata Herbst H.E.Andrewes det.” / “C. pseudoattenuata det. K. Kult 44”; ETHZ • 1 spec. [received with empty abdomen]; same data as for preceding; ETHZ • 1 ♂; “ Java ” / “Collection P. Dupuis” / “ Clivina ”; RBINS • 1 ♀; “ Java Koller”; RBINS • 1 ♀; “ Malang Java ” / “Collection P. Dupuis”; RBINS • 1 ♀; “ Java ” / “ Clivina ” / “ Collection P. Dupuis ”; RBINS • 1 ♀; “ Malang Java ” / “ Clivina attenuata Herbst H.E.Andrewes det. 21.XI.1925 ” / “C. pseudoattenuata m. det. K. Kult 44”; ETHZ • 1 ♀; “ Mt. Oengaran Java central. KOLLER” / “ Clivina ” / “ Collection P. Dupuis ”; RBINS • 1 ♀; “ Tjilatjap Java Drescher 15 XI 1926 ” / “ C. attenuata Hbst Andr. Dte. 23.11.33 ”; ETHZ .

Description

MEASUREMENTS. Holotype: body length 6.99 mm, width 2.03 mm; L/W of pronotum 0.94; L/W of elytra 1.79. Other material: body length 6.11–7.09 mm (ẍ: = 6.4 mm *), width 1.84–2.07 mm (ẍ: = 2.0 mm*), L/W of pronotum 0.91–0.95 (ẍ: = 0.94*), L/W of elytra 1.79–1.89 (ẍ: = 1.83*); (*n = 10).

COLOUR. Glossy. Piceous. Labrum, intermediate and hind leg fuscous, antenna pale fuscous. Supraantennal plates distinctly translucent-fuscous at the margin and slightly at middle.

HEAD. A third narrower than pronotum. Clypeus with central part distinctly bilobed anteriorly, lateral lobe distinctly projecting, lateral lobe and wing fused. Supraantennal plate nearly regularly convex laterally, laterally distinctly less projecting than eye;clypeus indistinctly reflexed margined, clypeal wing moderately and supraantennal plate distinctly reflexed margined. Supraantennal plate overlapping eye anteriorly by a fifth (dorsal view), with supraorbital keel extended nearly up to mid-eye level, separated from wing by right-angled notch. Clypeus transverse, moderately convex, separated from frons by slight step and furrow-like sulcus, sulcus in form like an inverted flat somewhat rounded V. Frons moderately convex, with two small but distinct arch-like impressions at middle, with supraorbital carina up to posterior eye-level. Clypeus smooth, frons with minute scattered punctures, separated from supraantennal plate and supraorbital carina by moderately deep and wide furrow, furrow interrupted at mid-eye level; with two supraorbital setigerous punctures, one situated at mid-eye level and one at gena level. Neck constriction missing. Eye globally projecting laterally. Main part of eye situated ventrally where it is globose. Due to the overlapping supraantennal plate and the gena, the eye looks in lateral view somewhat constricted. Gena distinct, moderately convex, covering a quarter of posterior eye in ventral view.Antenna of moderate length, reaching up to the middle of the pronotum, antennomeres five to ten elongate (L/W around 1.28). Labrum indistinctly trilobed anteriorly, with isodiametric reticulation, six setose. Mentum with isodiametric reticulation, with lobe flattened, shape of lobe asymmetric oval; distinctly carinate at middle, median tooth slightly dilated to apex, not as protruding anteriorly as lobe, flattened and obtusely rounded anteriorly.

PRONOTUM. Disk moderately convex in lateral view, in frontal view distinctly and regularly convex. Wider than long. Reflexed lateral margin smooth, nearly straight between middle and anterior setigerous puncture, widest behind middle; anterior angle slightly projecting, posterior angle missing; lateral channel moderately narrow, reticulated, completely and regularly rounded off at level posterior setigerous puncture and continuing as narrow channel up to base. Median line narrow, complete. Anterior transverse line moderately wide, formed by transverse punctures, joining median line, just not joining anterior margin. Surface glossy, basolaterally with impressed group of punctures of moderate size, with paramedian group of punctures, with few transverse wrinkles, whole disk covered with microscopic punctures. Channel at base relatively narrow. Proepisternum slightly visible in dorsal view.

ELYTRON. Disk flattened in anterior third in lateral view (in females slightly convex), distinctly convex in frontal view with increasing convexity towards lateral margin. Distinctly less than twice as long as wide, outline long-oval, lateral margin with straight part anteriorly to middle, with maximum width behind middle. Reflexed lateral margin smooth. Scutellar striole missing; setigerous tubercle at base of first stria, with small but distinct tubercle at base of second stria. Humeral tooth situated at base of fourth stria. Striae moderately deep, distinctly punctuate-striate, one to three free at base, one and two ending free at apex, three and four, and five and six joining apically. Intervals moderately convex, more convex laterally, interval eight carinate apically. Setigerous punctures missing on intervals. Surface of intervals glossy, interval eight with indistinct isodiametric reticulation covering about half of the width of the interval, in females interval seven with slight isodiametric reticulation directly at the humerus and the apical tip, in males smooth.

HIND WINGS. In the holotype, the alae are reduced by about 50%. Out of the other material, six specimens show fully developed alae, in the other material the alae are reduced up to a length and width of half of the elytron.

LOWER SURFACE. Proepisternum covered in lateral third with isodiametric reticulation and with distinct transverse wrinkles, medially with some punctures. Sternite of abdomen laterally with moderately large and dense punctures, smooth at middle but with microscopic punctures (in both sexes), apical segment with moderately large punctures, less dense basally; in females in addition with isodiametric reticulation laterally at the apical margin of the sternites three to five.

LEGS. Tibiae covered with isodiametric reticulation. Protibia with three spines of moderate length, the apical one distinctly curved ventrally, tibia not sulcate dorsally, movable spur robust, apically distinctly ventrally arcuate.

MALE GENITALIA (Fig. 31). Median lobe moderately massive, moderately arcuate at middle, with dorsal bulge in apical half, with complex, hollowed out and distorted apical lamella, with few microtrichia on the surface. Endophallus with around seven small teeth. Ventral paramere with velum-like apophysis, slightly distorted, apex with two setae. Dorsal paramere elongated sinus-like, with moderately developed apophysis, with two apical setae.

FEMALE GONOCOXITE AND EPIPLEURITE (Fig. 46). Gonocoxite less sclerotized, rhombic shaped, flattened dorsally and ventrally, with four long nematiform setae positioned at the median side, with one very long nematiform seta situated at the apical tip. Apical third pointed, curved and distorted in three dimensions. Epipleurite rectangular, slightly elongated, with distinctly widened angulated rod.

Variation

In some of the specimens, the anterior part of the pronotal lateral margin including the anterior angles is more or less translucent-fuscous. On the frons of the head, the arch-like impression varies in size. On the pronotum, the anterior transverse line is more or less formed by transverse punctures. Therefore, in some specimens it appears more as a line. The reticulation on interval eight of the elytron is in general indistinct. It consists of a small strip, which can reach half of the width of the interval in some of the specimens.

Sexual dimorphism

In females, the elytron is slightly convex in the anterior third in lateral view (flattened in males). In females, interval seven shows a slightly isodiametric reticulation directly at the humerus and at the apex, in males these parts are smooth.

Distribution

Known from the island of Java, Indonesia.

Figures

Figs 12–19. Sulciclivina Balkenohl, 2022, habitus, dorsal view. 12–13. S. striata striata (Putzeys, 1846). 12. ♂, lectotype, [specimen not cleaned], (OUMNH). 13. ♂ from Bandra, India (NHMUK). 14. S. striata kottea subsp. nov., ♀, holotype (NHMUK). 15. S. curvata sp. nov., ♂ from Java (ETHZ). 16–17. S. sulcigera (Putzeys, 1867). 16. ♂ from Nakhon, Thailand (NHMUK). 17. ♀ from Siem Reap, Cambodia, showing the reduced alae (RBINS). 18. S. karelkulti karelkulti sp. nov., ♀, holotype (MFNB). 19. S. karelkulti medanensis subsp. nov., ♀, holotype (SMNS).