Sulciclivina basiangusta, Balkenohl, 2023

Balkenohl, Michael, 2023, Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini), European Journal of Taxonomy 915 (1), pp. 1-57 : 19-21

Sulciclivina basiangusta sp. nov.

urn:lsid:zoobank.org:act:F940799A-2ECD-4785-B9C4-872F3F96BF56

Figs 9 , 29, 40, 53

Diagnosis

A medium-sized species with oval shape of the elytron and isodiametric reticulation present as a narrow strip on interval eight of the elytron close to the lateral channel. This is the only species of the genus with all intervals of the elytron distinctly convex and carinate apically. In addition, the base of the pronotum is extraordinarily narrow.

Etymology

The name refers to the extraordinarily small base of the pronotum and is combined from the Latin noun for base (ʻ basis ʼ) and the Latin adjective for narrow in the feminine form (ʻ angusta ʼ).

Type material

Holotype

INDIA • ♂; with labels and data: white, black printed “Megalaya 1976 Wittmer, Baroni U.” / “Mawphlang 15.5. 1850 m ”; NHMB.

Paratypes

INDIA • 1 ♂, 2 ♀♀; same data as for holotype, one of the females with additional label “ Clivina sagittaria Bates v. typ . Mus. Genova A. Casale det.”; NHMB, CMBB .

Description

MEASUREMENTS. Holotype: body length 6.91 mm, width 2.12 mm; L/W of pronotum 0.94; L/W of elytra 1.77. All material: body length 6.3–6.93 mm (ẍ: = 6.67 mm *), width 1.9–2.12 mm (ẍ: = 2.0 mm*), L/W of pronotum 0.91–0.97 (ẍ: = 0.94*), L/W of elytra 1.76–1.78 (ẍ: = 1.77*); (*n = 4).

COLOUR. Glossy. Piceous. Labrum, intermediate and hind leg fuscous, antenna and tarsomeres of front leg fuscous. Supraantennal plate at the margin translucent-fuscous.

HEAD. A third narrower than pronotum. Clypeus with central part distinctly bilobed anteriorly with the lobes bent dorsally; lateral lobe prominent, distinctly projecting anteriorly, clypeal wings indistinct, visible as slight convexity; supraantennal plate laterally regularly convex, laterally nearly as far projecting as eye; clypeus, wing and supraantennal plate reflexed margined. Supraantennal plate overlapping eye anteriorly by about a fifth (dorsal view), with the widened supraorbital keel extended up to mid-eye level, separated from wing by sharp notch. Clypeus transverse, with raising convexity posteriorly, separated from frons by distinct sharp carina and sulcus, carina and sulcus in form like an inverted flat V. Frons moderately convex, with two small impressions at middle, with supraorbital carina up to posterior eye-level. Clypeus smooth, frons with few transverse rugae at anterior eye-level, clypeus and frons separated from supraantennal plate and supraorbital carina by deep moderately wide furrow; with two supraorbital setigerous punctures at middle of the eye and end of gena-level. Neck constriction missing. Eye slightly convex in dorsal view, reduced, main part situated ventrally where it is convex. Gena enlarged, distinctly convex, covering half of posterior eye in ventral view. Due to the overlapping supraantennal plate and the genae, the eye is reduced to a strip in lateral view. Antenna moderately short, reaching just over middle of pronotum, antennomeres five to ten distinctly longer than wide (L/W around 1.3). Labrum indistinctly trilobed anteriorly, with isodiametric-irregular reticulation, six setose. Mentum with isodiametric reticulation, with lobe flattened, carinae on surface, shape of lobe asymmetric oval, with long median carina. Median tooth moderately wide, not as protruding anteriorly as lobe, flattened and rounded anteriorly.

PRONOTUM. Disk moderately convex in lateral and distinctly convex in frontal view. Slightly wider than long, subsquare. Reflexed lateral margin nearly smooth, indistinctly convex at middle, widest behind middle; anterior angle distinct, slightly projecting, posterior angle missing; lateral channel narrow, subcrenulated, completely and regularly rounded off at level of posterior setigerous puncture and continuing up to base. Median line deep, sharp, complete. Anterior transverse line consisting of longitudinal connected punctures, joining median line, not joining anterior margin. Surface glossy, covered densely with small punctures, with paralateral and paramedian group of punctures of moderate size, with few transverse wrinkles. Base distinctly marked, three times as wide as lateral channel, extraordinary narrow with less than a fifth of width of pronotum. Proepisternum distinctly visible in dorsal view.

ELYTRON. Shape oval. Disk moderately and completely convex in lateral view, distinctly convex in frontal view with increasing convexity to lateral margin. About 1.7 times as long as wide, with maximum width behind middle. Reflexed lateral margin smooth. Scutellar striole missing; setigerous tubercle at base of first stria, with distinct tubercle at base of third interval, basal declivity with isodiametric reticulation. Humeral tooth situated at base of fourth stria. All striae distinctly deepened, punctuate-striate, two and three finely punctate, one to three free at base, four reaching humerus, one and two ending free at apex, three and four, and five and six joining apically. Intervals distinctly convex, more convex laterally, all carinate apically. Setigerous punctures missing on intervals. Surface of intervals glossy, interval eight with small strip of reticulation close to lateral channel.

HIND WINGS. Alae reduced to a small, short rudiment (on the basis of the four type specimens).

LOWER SURFACE. Proepisternum covered in lateral half with distinct isodiametric reticulation and with some deep transverse wrinkles. All sternites of abdomen covered with moderately large punctures, with slightly less density at middle.

LEGS. Protibia covered with isodiametric reticulation, with three spines of moderate length, not sulcate dorsally, movable spur robust, in apical half moderately arcuate. Intermediate and hind tibia covered with irregular reticulation.

MALE GENITALIA (Fig. 29). Median lobe moderately slender, relatively short in comparison to body size, moderately arcuate at middle, with slight dorsal bulge, apex with moderately long, flattened, widened, and distorted spatula, with few microtrichia on the surface. Endophallus longitudinally folded. Ventral paramere with wide velum-like apophysis, somewhat distorted, apex with three setae. Dorsal paramere robust, attenuated up to apex, with four apical setae.

FEMALE GONOCOXITE AND EPIPLEURITE (Fig. 40). Gonocoxite triangle-shaped, flattened dorsally and ventrally, with four nematiform setae positioned at the median side, with one preapical long nematiform seta. Epipleurite semi-rectangular, with conspicuously widened angulated rod.

Variation

In addition to the sexual dimorphism, the following variations are observed: the sulcus between the carina of the clypeus and the frons is more or less widely impressed and in the impression irregularly structured. The lateral margin of the pronotum shows in two of the four specimen a few slight notches. The paramedian group of punctures in the basal half of the pronotum varies in the number of punctures from two to ten. Striae one and two of the elytron are joining at the apex in two of the paratypes. The intervals of the elytron are more or less carinate apically.

Sexual dimorphism

In males, the punctuation on the abdominal sternites is less dense at middle. Females show in addition isodiametric reticulation at the apical margins of the sternites.

Distribution

Known from the type locality Mawphlang, Meghalaya, in the North of India.

Figures

Figs 4–11. Sulciclivina Balkenohl, 2022, habitus, dorsal view. 4–5. S. attenuata attenuata (Herbst, 1806). 4. ♀, lectotype (MFNB). 5. ♂ from Swat (CMBB). 6. S. attenuata semireticulata subsp. nov., ♂, holotype (MNHN). 7. S. bhamoensis (Bates, 1892), ♀, paralectotype (RBINS). 8. S. mikirensis sp. nov., ♀, holotype (NHMB). 9. S. basiangusta sp. nov., ♂, holotype (NHMB). 10. S. coxisetosa sp. nov., ♀, holotype (NHMB). 11. S. splendida sp. nov., ♂, holotype (CADW).