Philibaetis Kaltenbach & Gattolliat, 2021

Kaltenbach, Thomas, Garces, Jhoana M. & Gattolliat, Jean-Luc, 2021, Philibaetis gen. nov., a new genus from the Philippines (Ephemeroptera, Baetidae), Deutsche Entomologische Zeitschrift 68 (1), pp. 1-20 : 1

publication ID	https://dx.doi.org/10.3897/dez.68.59462
publication LSID	lsid:zoobank.org:pub:458E5069-2CA2-436A-ABF5-BEFE6EBAC3F0
persistent identifier	https://treatment.plazi.org/id/5C936E67-64CD-43BC-AC21-35709769C2ED
taxon LSID	lsid:zoobank.org:act:5C936E67-64CD-43BC-AC21-35709769C2ED
treatment provided by	Deutsche Entomologische Zeitschrift by Pensoft
scientific name	Philibaetis Kaltenbach & Gattolliat
status	gen. nov.

Treatment

Philibaetis Kaltenbach & Gattolliat gen. nov. Figures 1 View Figure 1 , 2 View Figure 2 , 3 View Figure 3 , 4 View Figure 4 , 5 View Figure 5 , 6 View Figure 6 , 7 View Figure 7 , 8 View Figure 8 , 9 View Figure 9 , 10 View Figure 10 , 11 View Figure 11

Type species.

Philibaetis luzonensis ( Müller-Liebenau, 1982) comb. nov., by present designation.

Species included.

Philibaetis realonae ( Müller-Liebenau, 1982) comb. nov.

Diagnosis.

Larva. This new genus is distinguished by a combination of the following characters: A) body elongate and slender, head clearly hypognathous (Fig. 5 View Figure 5 ); B) bases of antennae not close to each other, without carina between them (Figs 5c View Figure 5 , 8c View Figure 8 ); C) labrum rectangular, on dorsal surface with submarginal row of long, simple setae (Fig. 1a View Figure 1 ); ventrally on lateral margins long, simple, spine-like setae, on anterolateral margins long, feathered setae and medially long, bifid setae and a partial, submarginal row of lanceolate setae (Figs 1b-f View Figure 1 , 6a, b View Figure 6 ); D) right mandible with blade-like incisor, stick-like, apically denticulate prostheca and dorsally mediolateral patch of long, spine-like setae (Fig. 1g, h, l View Figure 1 ); E) left mandible with blade-like incisor, slender prostheca apically denticulate and with comb-shaped structure; tuft of long, partly-branched setae at base of subtriangular process; dorsally mediolateral patch of long, spine-like setae (Fig. 1i-k, m View Figure 1 ); F) hypopharynx with a medial tuft of stout setae and anterolaterally two smaller tufts of stout setae (Fig. 2a View Figure 2 ); G) galea-lacinia with three denti-setae, distal denti-seta tooth-like and directed against canines, middle and proximal denti-setae slender, bifid and pectinate; maxillary palp 2-segmented, apex of segment II constricted (Fig. 2b View Figure 2 ); H) glossae basally broad, narrowing towards apex, shorter than paraglossae, apically and on both margins with long, spine-like setae; paraglossae truncate in distal part, one stout, bifurcate seta in posterolateral area; labial palps segment II without protuberance (Fig. 2d, e View Figure 2 ); I) fore femur apically with stout setae both on anterior and posterior side, femoral patch absent; claw robust and pointed, with one row of denticles and 2-3 long subapical setae on posterior side and one short, fine, subapical seta on anterior side (Figs 3a-c View Figure 3 , 11 View Figure 11 ); J) paraproct with patch of notched scales (Fig. 4e, f View Figure 4 ); K) protogonostyli under larval cuticle folded in the Labiobaetis type (Fig. 10 View Figure 10 ).

Imagines.

Unknown.

Etymology.

Philibaetis is an arbitrary combination of letters with allusion to the Philippines, where the genus occurs and to the genus Baetis . The latter is with reference to the superficial similarities between both genera and the fact that the species of Philibaetis gen. nov. were up to now assigned to Baetis . Gender is masculine.

Description.

Larva. (Figs 1 View Figure 1 - 11 View Figure 11 ).

Body. Elongate and slender, head hypognathous (Fig. 5 View Figure 5 ).

Head. Antenna (Figs 3d View Figure 3 , 5c View Figure 5 , 8c View Figure 8 ). More than 2 × longer than head length, bases of antennae not close to each other, without carina between them (Figs 5c View Figure 5 , 8c View Figure 8 ).

Labrum (Figs 1a-f View Figure 1 , 6a, b View Figure 6 ). Rectangular, wider than long; on dorsal surface with submarginal row of long, simple setae; ventrally on lateral margin long, simple, spine-like setae, on anterolateral margin long, feathered setae and medially long, bifid setae and a partial, submarginal row of lanceolate setae.

Right mandible (Fig. 1g, h, l View Figure 1 ). Incisor and kinetodontium fused, incisor blade-like, kinetodontium with denticles; inner margin of innermost denticle of kinetodontium without row of thin setae; prostheca stick-like, apically denticulate; dorsally with mediolateral patch of long, spine-like setae.

Left mandible (Fig. 1i-k, m View Figure 1 ). Incisor and kinetodontium fused, incisor blade-like, kinetodontium with denticles; prostheca apically denticulate and with comb-shaped structure; with tuft of long, partly-branched setae at base of subtriangular process; dorsally with mediolateral patch of long, spine-like setae.

Hypopharynx (Fig. 2a View Figure 2 ). With medial tuft of stout setae and anterolaterally two smaller tufts of stout setae.

Maxilla (Fig. 2b, c View Figure 2 ). Apically with three stout canines and three denti-setae; distal denti-seta tooth-like, with opposite direction to canines, other denti-setae slender, bifid and pectinate; maxillary palp with two segments.

Labium (Fig. 2d, e View Figure 2 ). Glossae basally broad, narrowing towards apex, shorter than paraglossae; apically and on both margins with long, spine-like setae; paraglossae truncate in distal part, on apex with three rows of long, robust, distally-pectinate setae and one stout, bifurcate seta in posterolateral area; labial palps segment II without protuberance, dorsally with row of spine-like setae near outer lateral margin.

Thorax. Hind protoptera (Fig. 4g View Figure 4 ). Absent.

Foreleg (Fig. 3a-c View Figure 3 ). Femur with dense row of curved, spine-like setae on dorsal margin, on apex short, stout setae on anterior and posterior side; femoral patch absent; tibia with patellotibial suture, dorsal margin with row of medium, clavate setae; claw robust and pointed, with one row of denticles, 2-3 long subapical setae on posterior side and one short, thin subapical seta on anterior side.

Abdomen. Gills (Figs 4a-d View Figure 4 ). Seven pairs of gills on segments I-VII, dorsally orientated. Margin with minute denticles intercalating fine, simple setae.

Paraproct (Figs 4e, f View Figure 4 ). Without prolongation at posterior margin, with stout, marginal spines; surface with patch of notched scales.

Caudal filaments (Fig. 5a, d View Figure 5 ). Inner margin of cerci with long, thin setae; paracercus bilaterally with long, thin setae.

Imago.

Unknown.

Distribution

(Fig. 12 View Figure 12 ). Philippines: Luzon.

Redescription