Zygadenia longicoxa, Soriano & Delclòs, 2006

Soriano, Carmen & Delclòs, Xavier, 2006, New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family, Acta Palaeontologica Polonica 51 (1), pp. 185-200 : 192-193

Zygadenia longicoxa sp. nov.

Figs. 6C , 7B .

Type locality: Las Hoyas fossil site, La Cierva, Serranía de Cuenca , Cuenca Province, Spain .

Stratigraphic horizon: Second lithosome of finely laminated limestones of La Huérguina Formation, Lower Cretaceous, Barremian (Fregenal−Martínez and Meléndez 2000).

Derivation of the name: After long coxae of this species.

Material.— Holotype MCCM LH 928030 part and counterpart. Beetle lacking antennae and legs. Another specimen ( MCCM LH 23432 ), part and counterpart of a distorted beetle without part of the head, antennae and legs. Both specimens from Las Hoyas.

Diagnosis.—Differs from species with similar ratio between last and penultimate ventrite in bigger size. From Z. mongolicus Ponomarenko, 1994 differs in strongly longer metacoxae.

Description.—Measurements: length 18.1 mm; width 9.3 mm; elytra length 12.7 mm. Body quite large, subovoid and flattened. Head transverse, about 1.5 times as wide as long without mandibles, and strongly retracted in pronotum; genae and temples almost as long as eyes, occiput sloped; vertex with two longitudinal keels, which in turn bear tubercles behind eyes. Neck−like constriction weak. Pronotum widest at base and narrowing anteriorly, about 1.3 times as wide as long at posterior edge, and almost as wide as long at anterior edge; anterior angles distinct; posterior angles drawn out. Disc of pronotum with flat elevation divided by a deep longitudinal groove. Prosternum nearly quadrate, about as long as mesosternum, and longer than metasternum. Procoxae rounded and contiguous. Mesocoxae oval and contiguous. Metasternum with well−expressed transverse and longitudinal sutures; metepisterna comparatively narrow. Metacoxae comparatively long, about twice longer than wider. Abdomen with 5 visible ventrites with sharply marked relief, their posterior half elevated above anterior half; last ventrite about 2.8 times as long as previous one. Elytra about three times as long as wide, its apex is not drawn out into “tail”. Epipleura comparatively narrow, with a row of indistinct cells. Cells of elytra rounded, with approximately 25 cells forming a row. Abdomen covered with comparatively small tubercles, larger on last ventrite and depressed parts of preceding ventrites.

Figures

Fig. 6. Barremian (Early Cretaceous) ommatins Brochocoleus (Brochocoleini) and Zygadenia (Notocupedini), and cupedins Priacma (Priacmini) and Anaglyphites (Mesocupedini) from Spain. A. Zygadenia diazromerali sp. nov. Holotype MCCM LH 12001a, part, Las Hoyas, Cuenca Province. B. Zygadenia martinclosas sp. nov. Holotype IEI LC 2663a, part, La Cabrúa, Lleida Province. C. Zygadenia longicoxa sp. nov. Holotype MCCM LH 908030a, part, Las Hoyas, Cuenca Province, Lower Cretaceous. D. Zygadenia siniestri sp. nov. Holotype MCCM LH 23515a, part, Las Hoyas, Cuenca Province. E. Priacma sanzii sp. nov. Holotype MCCM LH 24524, part, Las Hoyas, Cuenca Province. F. Anaglyphites zherikhini sp. nov. Holotype MCCM LH 15122a, part, Las Hoyas, Cuenca Province. G.?Anaglyphites pluricavus sp. nov. Holotype IEI LP 97Ga, part, La Pedrera, Lleida Province. Scale bars 2 mm.

Fig. 7. Camera lucida drawings of Barremian (Early Cretaceos) ommatin Zygadenia. A. Zygadenia martinclosas sp. nov. Holotype IEI LC 2663a, part, La Cabrúa, Lleida Province, dorsal (A1) and ventral (A2) aspects. B. Zygadenia longicoxa sp. nov. Holotype MCCM LH 928030a, part, Las Hoyas, Cuenca Province, dorsal (B1) and ventral (B2) aspects. Scale bars 2 mm.