Zygadenia martinclosas, Soriano & Delclòs, 2006

Soriano, Carmen & Delclòs, Xavier, 2006, New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family, Acta Palaeontologica Polonica 51 (1), pp. 185-200 : 191

Zygadenia martinclosas sp. nov.

Figs. 6B , 7A .

Type locality: Quarry near Rubies, La Cabrúa fossil site, Sta. Maria de Meià , Sierra del Montsec, Lleida Province, Spain .

Stratigraphic horizon: Calcaires lithographiques à Plantes et Vértébres de la Pedrera de Rubies Fm., Barremian, Lower Cretaceous ( Martín−Closas and López−Morón 1995).

Derivation of the name: Named after the Spanish palaeobotanist Carles Martín−Closas.

Material.— Holotype IEI LC 2663 part and counterpart, from La Cabrúa fossil site. The exemplar is an impression, well preserved, lacking part of antennae and legs. MCCM LH 20151 from Las Hoyas fossil site, part and counterpart of a beetle without head. The deformation of the fossil, especially on prosternum, metasternum and final part of abdomen, make impossible to assign this specimen without any doubt to this species.

Diagnosis.—Differs from other species with similar ratio between ventrites and similar size ( Z. sinitzae Ponomarenko, 2000 from the Lower Cretaceous of Transbaikalia and Z. patulus Ponomarenko, 1985b from the Lower Cretaceous of West Siberia) in narrower pronotum and elytra, and wider epipleura.

Description.—Measurements: length 16.1 mm; width 9.1 mm; elytra length 10.8 mm. Body rather elongate and large. Head transverse, about 1.5 times as wide as long without mandibles, narrowed roundly before and after eyes; cheeks and temples shorter than eyes, occiput sloped. Scape and pedicel about two times longer as each of antennomeres 3–8. Cervical constriction weak. Pronotum transverse, and abruptly narrowed anteriorly, about 1.2 times as wide as long at anterior edge, and about 1.8 times as wide as long at posterior edge; anterior angles sharply distinct, with no incised margin; lateral sides of pronotum somewhat flattened; posterior angles distinct. Disc of pronotum with large rectangular elevation divided by a longitudinal furrow. Prosternum 1.7 times shorter than mesosternum, and nearly two times shorter than metasternum. Procoxae rounded and contiguous. Mesosternum with a deep medial fossa in its last half. Mesocoxae rounded and contiguous. Metasternum with well−expressed longitudinal and transverse sutures; metepisterna comparatively wide. Abdomen with 5 visible ventrites with sharply marked relief, last ventrite about 2.5 times as long as previous one. Elytra about 1.4 times as long as its combined width, with apex not drawn out into “tail”; epipleura comparatively wide with a row of distinct cells extending to apex. Cells of elytra rounded and not elongated at posterior edge of elytra, with approximately 25 cells forming a row. Legs moderately developed; mesofemora somewhat elongate, widened posteriorly from its 1/3 posterior edge. Abdomen with no distinct tubercles.

Figures

Fig. 6. Barremian (Early Cretaceous) ommatins Brochocoleus (Brochocoleini) and Zygadenia (Notocupedini), and cupedins Priacma (Priacmini) and Anaglyphites (Mesocupedini) from Spain. A. Zygadenia diazromerali sp. nov. Holotype MCCM LH 12001a, part, Las Hoyas, Cuenca Province. B. Zygadenia martinclosas sp. nov. Holotype IEI LC 2663a, part, La Cabrúa, Lleida Province. C. Zygadenia longicoxa sp. nov. Holotype MCCM LH 908030a, part, Las Hoyas, Cuenca Province, Lower Cretaceous. D. Zygadenia siniestri sp. nov. Holotype MCCM LH 23515a, part, Las Hoyas, Cuenca Province. E. Priacma sanzii sp. nov. Holotype MCCM LH 24524, part, Las Hoyas, Cuenca Province. F. Anaglyphites zherikhini sp. nov. Holotype MCCM LH 15122a, part, Las Hoyas, Cuenca Province. G.?Anaglyphites pluricavus sp. nov. Holotype IEI LP 97Ga, part, La Pedrera, Lleida Province. Scale bars 2 mm.

Fig. 7. Camera lucida drawings of Barremian (Early Cretaceos) ommatin Zygadenia. A. Zygadenia martinclosas sp. nov. Holotype IEI LC 2663a, part, La Cabrúa, Lleida Province, dorsal (A1) and ventral (A2) aspects. B. Zygadenia longicoxa sp. nov. Holotype MCCM LH 928030a, part, Las Hoyas, Cuenca Province, dorsal (B1) and ventral (B2) aspects. Scale bars 2 mm.