Zygadenia oculata, Soriano & Delclòs, 2006

Soriano, Carmen & Delclòs, Xavier, 2006, New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family, Acta Palaeontologica Polonica 51 (1), pp. 185-200 : 190

Zygadenia oculata sp. nov.

Figs. 2F , 4B .

Type locality: Quarry near Rubies, La Cabrúa fossil−site, Sta. Maria de Meià , Sierra del Montsec, Lleida Province, Spain .

Stratigraphic horizon: Calcaires lithographiques à Plantes et Vértébres de la Pedrera de Rubies Fm., Barremian, Lower Cretaceous ( Martín−Closas and López−Morón 1995).

Derivation of the name: Named after Latin oculus (eye). Material.— Holotype IEI LC 3675 , part and counterpart, impression of beetle lacking most part of antennae and legs.

Diagnosis.—From species with similar ratio between last ventrite and penultimate one differs in size and general aspect of body. From Z. sinitzae Ponomarenko, 2000 differs in wider pronotum and elytra, and presence of a row of tubercles in the outer edge of pronotal disc.

Description.—Measurements: length 9.7 mm, width 4.5 mm; length of abdomen 6.0 mm; elytra length 6.7 mm. Body small and fairly elongate. Head transverse, about twice wider than longer without mandibles; genae and temples longer than eyes. Eyes oblique, about two times wider than longer, covered in their posterior part by pronotum. Mandibles comparatively short, somewhat shorter than eyes. Neck−like constriction indistinct. Pronotum transverse, about 1.6 times as wide as long at posterior edge, narrowing anteriorly; anterior angles drawn out, posterior angles obtuse. Disc of pronotum with large elevation divided by longitudinal groove, and with rounded tubercles in its inner margins. Prosternum almost as wide as long, as long as metasternum, and quite shorter than mesosternum. Procoxae rounded and contiguous. Mesocoxae oval. Metasternum about 1.8 times as wide as long. Abdomen with 5 visible ventrites with pronounced relief, their posterior half elevated above anterior half; last ventrite about 2.4 times as long as previous one. Elytra about three times as long as wide, apex not drawn out into “tail”. Epipleura comparatively narrow, with a row of indistinct cells. Cells of elytra rounded, about 25 form a row. Abdomen quite evenly covered with small tubercles, larger on last ventrite and depressed parts of preceding ventrites.

Figures

Fig. 2. Barremian (Early Cretaceous) ommatins Tetraphalerus, Cionocoleus (both Ommatini), Brochocoleus (Brochocoleini), and Zygadenia (Notocupedini) from Spain. A. Tetraphalerus ponomarenkoi sp. nov. Holotype MCCM LH 78a, part, Las Hoyas, Cuenca Province. B. Tetraphalerus penalveri sp. nov. Holotype DEPGM LP 0058b, counterpart, La Pedrera, Lleida Province. C. Cionocoleus longicapitis sp. nov. Holotype IEI LP 0164G, La Pedrera, Lleida Province. D. Brochocoleus indibili sp. nov. Holotype IEI LC 4636a, part, La Cabrúa, Lleida Province. E. Zygadenia viridis sp. nov. Holotype MCCM LH 1805a, part, Las Hoyas, Cuenca Province. F. Zygadenia oculata sp. nov. Holotype IEI LC 3675a, part, La Cabrúa, Lleida Province. Scale bars 2 mm.

Fig. 4. Camera lucida drawings of Barremian (Early Cretaceous) ommatin Zygadenia from Spain. A. Zygadenia viridis sp. nov. Holotype MCCM LH 1805a, part, Las Hoyas, Cuenca Province, dorsal (A1) and ventral (A2) aspects. B. Zygadenia oculata sp. nov. Holotype IEI LC 3675a, part, La Cabrúa, Lleida Province, dorsal (B1) and ventral (B2) aspects. Scale bars 2 mm.