Amara (Curtonotus) goniodera Tschitscherine , 1895

Li, Yihang, Li, Haoyuan & Shi, Hongliang, 2024, Revision of the macropterous subgenus Curtonotus from east China, with the description of a new species (Carabidae, Zabrini, Amara), ZooKeys 1190, pp. 39-73 : 39

Amara (Curtonotus) goniodera Tschitscherine, 1895

Fig. 4

Amara (Curtonotus) goniodera Tschitschérine, 1895: 164 (type locality: Korea; holotype in TMB); Hieke 1990: 256.

Specimens examined.

2 males (CBJFU), China, Inner Mongolia, Genhe, Daxinganling Ecological Station , 50.8061°N, 121.5824°E, 726 m, 2018.08.28, Hongliang Shi leg. ; 1 male (CBJFU), China, Jilin, Baishan, Fusong County, west of Changsongling tunnel, 41.7789°N, 127.9400°E, 1577 m, 2019.08.09, Hongliang Shi & Yizhou Liu leg. ; 1 male (CBJFU), China, Jilin, Antu County, north slope of Changbaishan mt, the waterfall of Tianchi , 42.0373°N, 128.0544°E, 1959 m, 2019.08.06, Hongliang Shi & Yizhou Liu leg. ; 1 female (CLYH), China, Jilin, Baishan, Antu County, north slope of Changbaishan mt, the waterfall of Tianchi , 2023.06.14, Taoqi Wang leg.

Chinese common name.

宽瓣暗步甲.

Diagnosis.

Medium-sized species, BL = 10.8-12.2 mm; dorsum black, legs dark drown; head small, its width ~ 1/2 of pronotum maximum width; head with one supraorbital seta each side. Pronotum (Fig. 4B ) cordate, densely punctate at base, sparsely punctate at mid-anterior region; lateral margins shallowly sinuate before posterior angles; posterior angles acute or nearly rectangular, slightly laterally protruded. Elytra elongated, widest near middle; sides of abdominal sternites densely punctate. Male mesotibiae projection (Fig. 4C ) not so distinct as A. (C.) gansuensis ; the proximal denticle blunt, present near apical third of tibia, distal denticles blunt and smaller than the proximal one, present at the midpoint between the proximal one and tibial distal apex. Male genitalia with apical lamella (Fig. 4F ) long and straight, subpointed and narrowed at tip; gonocoxite 2 (Fig. 4G ) of ovipositor elongate, broadly expanded, ~ 1.4-1.6 × as long as wide, slightly attenuate to apex, apex narrowly rounded.

Comparison.

This species can be distinguished from the other four Chinese species belonging to this species group by having a much longer and somewhat pointed apical lamella of the median lobe. Compared with A. (C.) tumida and A. (C.) shinanensis , A. (C.) goniodera has wider and stouter gonocoxite; compared with A. (C.) gansuensis , A. (C.) goniodera , it has narrower pronotum with narrower lateral groove and shorter lateral marginal sinuation before posterior angles; compared with A. (C.) hyperborea , A. (C.) goniodera , A. (C.) goniodera has distinctly punctate abdominal sternites.

Distribution.

China (Gansu, Heilongjiang, Qinghai, Jilin, Shaanxi, Inner Mongolia*, Hebei*), North Korea, South Korea, Mongolia, Russia (Far East, east Siberia).

Figures

Figure 4. Amara (Curtonotus) goniodera A dorsal habitus, male (Zhangbei, Hebei) B pronotum posterior angle C male mesotibia D lateral view of aedeagus E right paramere F apical lamella G female gonocoxite (Zhangbei, Hebei). Scale bars: 5 mm (A); 1 mm (B-E); 0.5 mm (F, G).