Manota rara, Jaschhof & Hippa, 2005

Jaschhof, Mathias & Hippa, Heikki, 2005, The genus Manota in Costa Rica (Diptera: Mycetophilidae), Zootaxa 1011 (1), pp. 1-54 : 10-12

Manota rara sp. n. ( Fig. 3C )

Description. Large species.

Male. Head. Antenna with fourth flagellomere 2.4 times as long as wide. Maxillary palpus with third palpomere bearing 3 curved sensilla; fourth palpomere with a parasegment apically.

Thorax. Anterior basalare bare. Laterotergite setose. Episternum 3 with ventral portion setose. Wing. Length 2.2 mm. Membrane non­setose. Sc distally of h bare. Junction of CuA2 into CuA1 incomplete.

Terminalia ( Fig. 3C ). Sternite 9 large, with posterior and most of lateral margins distinct; posterior margin straight, situated beyond mid length of gonocoxites; with setae larger than those on gonocoxites. Ventro­lateral lobes of gonocoxites small, subrectangular. Disto­lateral lobes of gonocoxites small and only dorsally developed. Gonocoxites in position I+II with a circle of some 10–15 short setae, some setae very thick and blunt, others stiff and pointed; in position III with 2 flat megasetae, each two­branched with one branch whip­like and the other branch rounded; with 1 large, posteriorly directed seta displaced from position IV into dorso­subapical position. Gonostyli elongate, bilobed; posterior lobe large, rounded, with numerous large setae; inwardly directed lobe smaller, subtriangular in ventral view, with some 10 setae marginally, obviously with a dorsal extension bearing 2 short setae. Parastylar lobes small, largely hidden behind gonostyli; each lobe with 2–3 setae apico­posteriorly. Parameral apodemes widely spread. Tegmen constricted in distal half; margin of ventro­lateral lobes distinct. Cerci unusually wide, longer than hypoproct. Hypoproct with short setulae disto­laterally.

Diagnostic characters. Among the species with long antennae and setose laterotergites, it has terminalia with very short disto­lateral lobes of the gonocoxites; elongate, bilobed gonostyli; unusually wide cerci; and gonocoxites with a characteristic circle of 10­ 15 short, stiff setae in position I+II.

Etymology. From Latin, rarus, scarce, referring to the evident scarcity of this species.

Holotype. Male, Costa Rica, Limón province, Hitoy Cerere Biological Reserve, Sendero Espavel , 560 m, in tall secondary rain­forest, 11 March–1 April 2003, by Malaise trap, E. Rojas, B. Gamboa & W. Arana ( INBio sample #73628).

Figures

FIGURE 3. Male terminalia of Manota spp. A: M. defecta Will. (lectotype), ventral view. B: M. defecta Will. (lectotype), detail of gonocoxites showing vestiture in positions I and II, ventral view. C: M. rara sp. n. (holotype), ventral view. Scale = 0.1 mm.