Phoenicocoris opacus (Reuter)

SCHWARTZ, MICHAEL D. & STONEDAHL, GARY M., 2004, Revision of Phoenicocoris Reuter with Descriptions of Three New Species from North America and a New Genus from Japan (Heteroptera: Miridae: Phylinae), American Museum Novitates 3464 (1), pp. 1-56 : 27-37

Phoenicocoris opacus (Reuter)

Psallus opacus Reuter, 1906: 72 (n.sp.); Schuh, 1995: 411 (catalog); Kerzhner, 1997: 247 (lectotype designation).

Phoenicocoris opacus: Kerzhner, 1997: 247 (n.comb.); Kerzhner and Josifov, 1999: 388 (catalog).

REDESCRIPTION: (N = 13, 5♀) GENERAL ASPECT: total length male 3.20, female 3.00 (2.90–3.20); length to cuneal fracture male 2.25, female 2.20 (2.15–2.40); overall coloration dirty yellow to pale brown; cuneus and apex of exocorium slightly darker than remainder of hemelytra; dorsum with narrow, flattened, and apically pointed silvery scalelike setae mixed with suberect dark brown simple setae; thoracic pleura, except propleura, and venter without scalelike setae. HEAD: width male 0.68, female 0.67 (0.65– 0.68); vertex width male 0.33, female 0.38; anterocular length male 0.20, female 0.22 (0.20–0.23); vertex usually with pale area medially; antennae measurement male 0.25: 0.88: 0.48: 0.30, female 0.26 (0.25–0.26): 0.84 (0.80 –0.88): 0.53 (0.48–0.58): 0.35(0.32–0.38); labium always reaching to, usually slightly surpassing apices of hind coxae; labial length male 0.11, female 0.13 (0.12–0.13). THORAX: width male 1.0, female 1.04 (0.98–1.08); length male 0.43, female 0.45 (0.43–0.48); dirty yellow to brown. HEMELYTRA: maximum width male 1.25, female 1.38 (1.30–1.45). LEGS: pale yellow; femora covered with distinct dark brown dots of various sizes apically; tibia with large dark brown dots at bases of dark spines.

DISCUSSION: We have not personably examined specimens of this taxon; however, Kerzhner (1997), when designating the lectotype of opacus , tentatively placed it in Phoenicocoris . The new combination was based on one teneral male and 10 female syntypes. The vesica of opcaus has two unequal apical processes. Kerzhner (1997) suggested that some external characters of P. opacus are similar to P. kyushuensis , while also noting that its eyes were much smaller and that its pale femora had dark spots. Linnavuori (1961), when describing kyushuensis , contrasted his new species with opacus stating that in the latter the vertex is 2 to 2.5 times as wide as the dorsal eye width and that the labium extends to the hind coxae. Herein we erect a new genus for P. kyushuensis based, in part, on the equal length vesical processes of the two included species. We maintain the generic assignment of opacus in Phoenicocoris and do not transfer it to our new genus because the vesical structure, eye size, labial length, and femoral spots are within the range of variation we attribute to Phoenicocoris . Current collection records of opacus indicate that its distribution is limited to Khunchyao Pass, Sichuan province in the southwest territory of China. The host plants of this species are unknown.

SPECIMEN INFORMATION: CHINA: Sichuan: les [=forest], sev[ernyi]. skl[on]. [=northern side), Khunchyao perev[al]. (=pass), 11.VIII.1893, Potanin, lectotype 3 ( ZISP); Tatszinlu, 21.V.1893 – 22.VI.1893, Potanin 10 ♀ paralectotypes ( ZISP).

Phoenicocoris pallidicornis , new species

Figures 1P, 3 , 16 , 24

HOLOTYPE: Male. ‘‘ Can [ada]. W. Ontario

Black Sturg[eon] Lake 1–15.VIII.1956 [H.]

Lindberg.’’ Deposited in the Canadian National Collection of Insects, Arachnids, and Nematodes.

DIAGNOSIS: Recognized by the bicolored dorsum with the head, scutellum, and most of pronotum dark brown, posterior portion of pronotum and hemelytra yellowish brown; short anteocular portion of head; wide vertex; and long, roughened gonopore sclerite (fig. 24). Distribution of scalelike setae as in australis , claricornis , dissimilis , and longirostris , but the contrasting dorsal coloration will distinguish pallidicornis from these species with uniform dorsal coloration.

DESCRIPTION: GENERAL ASPECT: broad, relatively short; total length male 3.00 (2.75– 3.15), female 2.92 (2.80–3.10); length to cuneal fracture male 2.10 (1.95–2.20), female 2.11 (2.00–2.18); coloration with head, anterior portion pronotum, including calli, mesoscutum, and scutellum dark brown to brownish black; posterior portion of pronotum and hemelytra pale yellowish brown; antennae, apices of femora, tibiae, and tarsi concolorous with hemelytra; ventral aspect concolorous with head; dorsum, excluding vertex, with moderately to densely distributed, broad, flattened, apically serrate, white scalelike setae and moderately densely distributed, suberect to reclining, black simple setae; thoracic pleura and venter without scalelike setae. HEAD: width male 0.77 (0.76–0.78), female 0.80 (0.77–0.82); vertex width male 0.39 (0.38–0.40), female 0.41 (0.39–0.43); frons and vertex broadly curved in lateral view; width of head and vertex wide; region anterior to antennal insertion short, anteocular length male 0.21 (0.19– 0.22), female 0.22 (0.19–0.23); eyes relatively small, about 76% of head height, ventral margin of antennal insertion ventral to ventral margin of eye; antennal measurements male 0.18 (0.18–0.19): 0.60 (0.58–0.61): 0.38 (0.34–0.41): 0.28 (0.25–0.30), female 0.18 (0.18–0.19): 0.61 (0.54 –0.63): 0.38 (0.35–0.41): 0.29 (0.25–0.31); antennal segment 2 is 20% shorter than width of head across eyes; labium yellowish brown, reaching middle of hind coxa; labial length male 1.01 (0.94–1.06), female 1.04 (1.00–1.09); length of segment 4 male 0.28 (0.26–0.30), female 0.28 (0.26–0.29). THORAX: width male 1.00 (0.94–1.03), female 1.01 (0.95– 1.06); length male 0.43 (0.41–0.45), female 0.44 (0.42–0.45); mesoscutum moderately exposed. HEMELYTRA: maximum width male 1.27 (1.23–1.38), female 1.35 (1.28– 1.40); lateral margin gently arcuate. LEGS: with diffuse dark brown spots at bases of long tibial spines.

BIOLOGY: Breeds on jack pine ( Pinus banksiana Lambert )

DISTRIBUTION: From disparate localities in New Brunswick, Ontario, western Québec, eastern Manitoba, and central Saskatchewan.

PARATYPES: CANADA: Manitoba: Falcon Lake , 14.VII.1972, L. A. Kelton, P. banksiana , 43, 10♀ ( CNC) . New Brunswick: Kouchibouguac National Park , 24.VII.1972, D. J. E. Brown, jack pine, [codes 5736P, 5764 R], 3♀ ( CNC) . Ontario: Black Sturgeon Lake , 1–15.VIII.1956, H. Lindberg, 13, 24♀ ( CNC) ; MacKey , 27.VII.1962, Kelton, Thorpe, P. banksiana , 1♀ ( CNC) ; Sibley Park , 1.VIII.1956, H. Lindberg, 4♀ ( CNC) ; Sioux Narrows , 8.VIII.1960, Kelton, Whitney, 2♀ ( CNC) ; Stittsville , 12.VII.1979, D. J. E. Brown, 23, 2♀ ( CNC) . Québec: Kazabazua , 15.VIII.1961, D. J. E. Brown, P. banksiana , 1♀ ( CNC) ; Laniel , 10–11.VII.1963, L. A. Kelton, 13, 6♀ ( CNC) . Saskatchewan: Torch River , 11.VII.1950, L. A. Konotopetz, P. banksiana , 33, 8♀ ( CNC) ; White Fox , 25.VII.1950, L. A. Konotopetz, Potentilla fruticosa L., 1♀ ( CNC) .

Phoenicocoris ponderosae , new species

Figures 1Q, 3 , 17 , 24

HOLOTYPE: Male. ‘‘[ USA.,] COLO[rado]., Park Co. 3 mi S. of Guffey VII–30–1983 D. A.& J. T. Polhemus , Pinus ponderosa (Ponderosa Pine) .’’ Deposited in the American Museum of Natural History.

DIAGNOSIS: Easily recognized by the large body size (2.90–3.65 mm); absence of scalelike setae on the vertex, pronotum, thoracic pleura, and abdominal sternum; and base of gonopore sclerite with conspicuous spinules (fig. 24). Further distinguished from modestus and nevadensis , which both have similarly distributed scalelike setae, by the much longer labium and large overall size, respectively.

DESCRIPTION: (N = 63) GENERAL AS­ PECT: broad, moderately elongate; total length male 3.31 (2.90–3.60), female 3.43 (3.15–3.65); length to cuneal fracture male 2.51 (2.38–2.70), female 2.55 (2.45–2.60); coloration dark brownish black; antennae, apices of femora, tibiae, and tarsi pale to dusky yellow; apex of tarsal segment 3 and claws darkened; mesoscutum, scutellum, and hemelytra with moderately to densely distributed, broad, flattened, apically serrate, white scalelike setae and densely distributed, suberect to reclining, brown simple setae; thoracic pleura and venter without scalelike setae. HEAD: width male 0.80 (0.79–0.81), female 0.83 (0.80–0.89); vertex width male 0.35 (0.33–0.36), female 0.38 (0.36–0.41); region anterior to antennal insertion elongate, anteocular length male 0.26 (0.24–0.28), female 0.27 (0.26–0.28); eyes relatively small, about 79% of head height, ventral margin of antennal insertion level with ventral margin of eye; antennal measurements male 0.20 (0.19–0.21): 0.74 (0.73–0.76): 0.46 (0.44– 0.49): 0.34 (0.31–0.37), female 0.20 (0.19– 0.20): 0.76 (0.69–0.82): 0.49 (0.44–0.52): 0.33 (0.28–0.35); antennal segment 2 is 7% shorter than head width; labium yellowish brown, reaching sternite 5 to base of genital segment in males and to base of ovipositor to middle of ovipositor in females; labial length male 1.79 (1.70–1.86), female 1.85 (1.76–1.93); length of segment 4 male 0.54 (0.51–0.56), female 0.56 (0.54–0.59). THO­ RAX: width male 1.16 (1.11–1.23), female 1.21 (1.15–1.30); length male 0.56 (0.54– 0.60), female 0.59 (0.56–0.64); mesoscutum broadly exposed. HEMELYTRA: maximum width male 1.44 (1.35–1.53), female 1.49 (1.41–1.58); lateral margin gently curved.

BIOLOGY: Breeds on ponderosa or yellow pine ( Pinus ponderosa Douglas ex Lawson and C. Lawson )

DISTRIBUTION: Central Rocky Mountains from Colorado southwest to Chiricahua Mountains , Arizona .

PARATYPES: USA: Arizona: Cochise Co. : Rustler Park, Chiricahua Mountains, 7, 8.VII.1968, L. A. Kelton, Pinus ponderosa , 33, 5♀ (CNC). Coconino Co.: Happy Jack, Coconino National Forest, 7600 ft, 13.VII.1968, L. A. Kelton, P. ponderosa , 1♀ (CNC). Colorado: Chaffee Co.: 5 mi W of Buena Vista, 25.VII.1982, J. T. Polhemus, 13 (JTP). La Plata Co.: Durango, 19.VII.1968, L. A. Kelton, P. ponderosa , 3♀ (CNC). Park Co.: 3 mi S of Guffey, 30.VII.1983, D. A., J. T. Polhemus, P. ponderosa , 13, 4♀ (JTP).

Phoenicocoris rostratus (Knight)

Figures 1U, 3 , 18 , 24

Lepidopsallus rostratus Knight, 1923: 470 (n.sp.).

Phoenicocoris rostratus: Stonedahl, 1990: 61 (n.comb.); Schuh, 1995: 375 (catalog).

Lepidopsallus olseni Knight, 1923: 473 (n.sp.). NEW SYNONYM.

Phoenicocoris olseni: Stonedahl, 1990: 61 (n.comb.); Schuh, 1995: 375 (catalog).

Lepidopsallus hesperus Knight, 1968: 53 (n.sp.). NEW SYNONYM.

Phoenicocoris hesperus: Stonedahl, 1990: 61 (n.comb.); Schuh, 1995: 374 (catalog).

DIAGNOSIS: Recognized by the broad, apically serrate scalelike setae on the thoracic pleura and lateral margins of abdominal sternum; antennal segment 2 yellow or yellowish brown, cylindrical, length less than or subequal to width of head across eyes; and structure of the male vesica (fig. 24), especially the absence of spinules on the distal edge of the secondary gonopore. Further distinguished from ponderosae and longirostris in the western montane United States by the shorter anteocular region of the head and shorter labium, especially segment 4.

REDESCRIPTION: GENERAL ASPECT: weakly elongate; total length male 2.86 (2.75–3.03), female 2.77 (2.58–2.99); length to cuneal fracture male 2.05 (1.98–2.15), female 2.07 (1.90 –2.30); coloration dark brown to nearly black; antennal segment 2– 4, apices of femora, tibiae, and tarsi pale yellow to dusky brownish yellow; tibiae basally and antennal segments 3 and 4 sometimes darker brown; dorsum, thoracic pleura, and lateral margins of abdominal sternum with moderately to densely distributed, broad, flattened, apically serrate, white scalelike setae and densely distributed, relatively long, suberect to reclining, black simple setae. HEAD: width male 0.73 (0.70–0.75), female 0.76 (0.73–0.79); vertex width male 0.33 (0.33– 0.34), female 0.35 (0.33–0.36); anteocular region relatively long, anteocular length male 0.20 (0.19–0.22), female 0.21 (0.19–0.24); eyes moderately large, occupying about 80% of head height, ventral margin of antennal insertion slightly ventral to ventral margin of eye; antennal measurements male 0.18 (0.18–0.19): 0.72 (0.70–0.75): 0.44 (0.41– 0.46): 0.30 (0.28–0.31), female 0.19 (0.18– 0.20): 0.69 (0.64–0.75): 0.45 (0.40–0.50): 0.32 (0.30–0.35); antennal segment 2 is 6% shorter than or equal to width of head across eyes; labium brown, reaching to just beyond hind coxae, or extending to, at most, middle of venter; labial length male 1.38 (1.33– 1.53), female 1.40 (1.34–1.45); length of segment 4 male 0.39 (0.35–0.43), female 0.41 (0.38–0.43). THORAX: maximum width male 1.00 (0.98–1.03), female 1.03 (1.00–1.10); length male 0.47 (0.41–0.52), female 0.47 (0.43–0.50); mesoscutum broadly exposed. HEMELYTRA: maximum width male 1.28 (1.23–1.35), female 1.32 (1.23– 1.40); slightly curved laterally.

BIOLOGY: Breeds on pines, with Pinus banksiana Lambert and P. sylvestris Linnaeus the predominate hosts in the east and P. contorta Douglas ex Loudon (including var. latifolia Engelmann Critchfield ) apparently the preferred host in the west. Also collected on the following conifers: shortleaf pine ( P. echinata Miller ), red pine ( P. resinosa Aiton ), pitch pine ( P. rigida Miller ), Scots pine ( P. sylvestris Linnaeus ), balsam fir ( Abies balsamea (Linnaeus) Miller ), Norway spruce ( Picea abies (Linnaeus) Karsten ), white spruce ( P. glauca (Moench) Voss ), and European larch ( Larix decidua Miller ).

DISTRIBUTION: Southern Yukon Territory and British Columbia , east to Nova Scotia. Widely distributed in montane regions of the western United States from Montana south to northern Arizona and California, east across the northern tier of states to New Hampshire and south to West Virginia.

DISCUSSION: Knight (1923) described both olseni and rostratus in Lepidopsallus Knight , based on single females from Pigeon Cove, Massachusetts and Elkhorn Creek, Minnesota, respectively. Knight’s (1923) key to Lepidopsallus , which in our taxonomy included three Phoenicocoris species ( claricornis , minusculus , and rostratus ) and two Atractotomus species ( atricolor and rubidus ), distinguished his new species based on the length of the labium and antennal segment 2. The description for olseni ( Knight, 1923) stated that antennal segment 2 is long (surpassing the length of the pronotum) and slen­ der, the venter is covered with scalelike setae, and that these setae are not apically truncate. Based on these features olseni cannot be confused with claricornis or minusculus ; the former does not have scalelike setae on the venter, and the antennal segment 2 of the latter is fusiform and has truncate scalelike setae. Antennal segment 2 surpassing the length of the pronotum will distinguish all species of Phoenicocoris from both A. atricolor and A. rubidus . Only the ‘‘short’’ labium distinguishes olseni from rostratus . Knight (1923) observed that the labium of olseni did not extend beyond the hind coxae; however, as he also noted, and we can verify (upon seeing the holotype), this structure was imbedded in glue and impossible to accurately measure. Our examination of many females of rostratus revealed that the length of the labium is variable, reaching from the apices of the hind coxae to as far as sternite 5. Determining the true labial length in specimens in which it is either bent or obscured by glue can be difficult and usually provides a shortlength measurement. Based on these arguments we surmise that Knight was examining a typical female of rostratus and we therefore propose L. olseni as a junior synonym of L. rostratus .

Knight (1968) described L. hesperus from California, Idaho, and Wyoming, apparently without considering the eastern North American species of his genus. Specimens from throughout the eastern portion of the range of P. rostratus will reach, without trouble, the couplet for hesperus in Knight’s key (1968). Western specimens will likewise reach the couplet for rostratus in Knight’s key (1923). Knight’s characterizations of both nominal species, hesperus and rostratus , adequately describe specimens from across North America and examination of paratypes of the former species and the holotype of the latter species shows that they fit comfortably within our concept of rostratus . As noted above, the majority of the eastern specimens examined were taken on Pinus banksiana whereas P. contorta was the predominate host of western specimens. These closely related pine species are shade­intolerant, common on well­drained sites, have indistinguishable diploxylon fossil pollen, and hybridize in west­central Alberta (Mac­ Donald, et al. 1998). No morphological characters allowed for the differentiation of Phoenicocoris hesperus specimens based on host. We found no variation in the form for either the apices of the vesical strap or the secondary gonopore in vesical preparations of P. rostratus from across North America. The variation in the vesicae documented in figure 24 (compare rostratus 1 and 2) is due to a slight rotation of the preparations. We therefore treat L. hesperus as a junior synonym of L. rostratus .

SPECIMENS EXAMINED: CANADA: Alberta: Brooks, Special Crops and Horticultural Research Centre, 17.VII.1990, M. D. Schwartz, Pinus sylvestris , 3♀ (CNC); Canmore, 27.VIII.1955, A. R. Brooks, Picea glauca , 1♀ (CNC); Coal Valley, 31.VIII.1970, L. A. Kelton, Pinus contorta , 2♀ (CNC); Coleman, 25 mi N of, 14.VII.1988, R. Wharton, 13 (TAM); Cypress Hills Provincial Park: Elkwater Lake, 20.VII.1956, O. Peck, P. contorta , 13 (CNC); Elkwater Park, 21, 26.VII, 14.VIII.1952, A. R. Brooks, L. A. Konotopetz, P. contorta , 33, 1♀ (CNC); Top Road, 9.6 km E of jct Rte 41 and Top Road, 4700 ft, 15.VII.1990, M. D. Schwartz, P. contorta 13, 2♀ (CNC). Eisenhower Junction, 5.VIII.1970, L. A. Kelton, P. contorta , 13, 1♀ (CNC); Fisher Creek Forest Station, 26.VII.1973, L. A. Kelton, 23 (CNC); Grand Prairie, 25.VII.1961, A. R. Brooks, 2♀ (CNC); Kananaskis Highway, 20.VII.1974, 25.VII.1973, L. A. Kelton, P. contorta , 23, 1♀ (CNC). Peace River, 18.VI.1961, A. R. Brooks, 5♀ (CNC); Waterton Lakes, 22.VII.1923, J. McDunnough, 13 (CNC); Waterton National Park, 4–6.VII.1970, 26.VII.1970, L. A. Kelton, P. contorta , 73, 2♀ (CNC). British Columbia : Arrowhead, 29.VII.1947, P. contorta var. latifolia, FIDS # BC47–1383, 1♀ (CNC); Aspen Grove, 25.VII.1970, L. A. Kelton, P. contorta , 63, 20♀ (CNC); Blackwall, Manning Provincial Park, 23.VII.1970, L. A. Kelton, 23 (CNC); Christina Lake, 12.VII.1970, L. A. Kelton, P. contorta , 43, 7♀ (CNC); Cranbrook, 23.VII.1959, L. A. Kelton, P. ponderosa , 23, 6♀ (CNC); Crowsnest Provincial Park, 4 km W of Crows Nest Pass, 1.IX.1993, M. D. Schwartz, P. contorta , 13 1♀ (CNC); Duncan, 16.VII.1959, L. A. Kelton, 1♀ (CNC); Elko, E Kootenay, 9.VII.1949, H. B. Leech, 1♀ (CAS); Fernie, L. A. Kelton: 7.VII.1970, P. contorta , 53, 4♀ (CNC); 23.VII.1959, 1♀ (CNC). Fruitvale, 11.VII.1970, L. A. Kelton, 13 (CNC); Gray Creek, 22.VII.1959, L. A. Kelton, 1♀ (CNC); Hedley, 22.VII.1970, L. A. Kelton, P. contorta , 73, 5♀ (CNC); Jaffray, 8.VII.1970, L. A. Kelton, P. contorta , 63, 5♀ (CNC); Kaslo, 10.VII.1970, L. A. Kelton, P. contorta , 23, 4♀ (CNC); Lumby, 13.VIII.1975, L. A. Kelton, 13, 2♀ (CNC); Moyie, 9.VII.1970, L. A. Kelton, 33, 2♀ (CNC); Osoyoos, 21.VII.1970, L. A. Kelton, 53 (CNC). Princeton, N. H. Anderson, P. ponderosa : 30 mi W of, 26.VII.1957, # 288, 13 (CNC); 15 mi W of, 31.VII.1957, # 290, 1♀ (CNC). Rossland, 11.VII.1970, L. A. Kelton, 2♀ (CNC); Sicamous, 17.VII.1970, L. A. Kelton, 2♀ (CNC); Summerland, 2–11.VII.1974, L. A. Kelton, P. sylvestris , 73, 6♀ (CNC); Terrace: 27.VII.1960, W. R. Richards, 13, 1♀ (CNC); 220 ft, B. S. Heming: 29.VII.1960, 1♀ (CNC); 31.VII.1960, 13 (CNC). Yahk, 22.VII.1959, L. A. Kelton, P. contorta , 153, 14♀ (CNC); Yoho National Park, 16.VIII.1970, L. A. Kelton, 13 (CNC). Manitoba: Carberry, W of, 1.0 km SE of jct. Hwy 1 and Rte 351 on Rte 351, 8.VII.1990, M. D. Schwartz, P. banksiana , 1♀ (CNC); Falcon Lake, 1, 5, 12.VII.1972, 6.VIII.1975, 14.VIII.1972, L. A. Kelton, P. banksiana , 93, 18♀ (CNC); Riding Mountain National Park, 20.VII.1972, L. A. Kelton, 1♀ (CNC); Sandilands Provincial Park, just S of Hwy 1, 19.VII.1990, M. D. Schwartz, P. banksiana , 23, 2♀ (CNC); West Hawk Lake, 28.VII.1972, L. A. Kelton, P. banksiana , 43, 5♀ (CNC); Victoria Beach, 18.VIII.1953, A. R. Brooks, L. A. Kelton, 2♀ (CNC). New Brunswick: Kouchibouguac National Park, 12, 22, 24, 28.VII.1977, D. J. E. Brown, D. B. Lyons, P. banksiana , 53, 12♀ (CNC); Penobsquis, 6.VIII.1978, L. A. Kelton, P. banksiana , 33, 1♀ (CNC); Sussex, 2.VIII.1966, L. A. Kelton, P. banksiana , 6♀ (CNC). Nova Scotia: Coldbrook, 10.VII.1966, L. A. Kelton, 1♀ (CNC); Halifax, 22.VII.1975, 7–8.VIII.1976, L. A. Kelton, P. sylvestris , 23, 18♀ (CNC); Kentville, 15– 17.VII.1966, L. A. Kelton, P. sylvestris , 7♀ (CNC); Middleton, 9.VIII.1966, P. resinosa , 1♀ (CNC); Springhill, 12.VIII.1966, L. A. Kelton, P. banksiana , 1♀ (CNC). Ontario: Almonte, 4 mi SW of, alvore, 26.VI.1990, M. D. Schwartz, P. banksiana , 73, 3♀ (CNC); Black Hawk, 3.VIII.1960, Kelton, Whitney, P. banksiana , 3♀ (CNC); Black Sturgeon Lake, 1–15.VIII.1956, H. Lindberg, 2♀ (CNC); Dryden, 12.VIII.1960, Kelton, Whitney, Salix sp. , 1♀ (CNC); Ignace, 5 km W of, on Hwy 17 roadside, M. D. Schwartz, P. banksiana , 23, 1♀ (CNC); MacKey, 27.VII.1962, Kelton, Thorpe, P. banksiana , 103, 26♀ (CNC); Myers Road, 2 km E of Vermillion Bay, gravel pit, 4.VII.1990., M. D. Schwartz, P. banksiana , 13, 1♀ (CNC); Nepean, Piney forest, Lafontaine house, M. D. Schwartz, MV light, 27.VII.1991, 13 (CNC); Nestor Falls, 20.VII.1978, 7– 8.VIII.1960, Kelton, Whitney, P. banksiana , 13, 14♀ (CNC). Ottawa, Central Experimental Farm, M. D. Schwartz: ca. K. W. Neatby Building, 7, 16.VI.1999, P. banksiana , P. rigida , 123, 9♀ (CNC); 19.VI.1991, P. sylvestris , ( Larix decidua 23), 1063, 179♀ (CNC); 15.VII.1994, P. sylvestris , 43, 12♀ (CNC). Shawanaga, 26.VII.1962, L. A. Kelton, G. Thorpe, P. banksiana , 23, 18♀ (CNC); Sioux Narrows, 8.VIII.1960, Kelton, Whitney, 3♀ (CNC); Stittsville, 12.VII.1979, D. J. E. Brown, 13, 3♀ (CNC); Tobermory, 1.VII.1962, G. Thorpe, P. banksiana , 33 (CNC); Vermillion Bay, 5–9.VII.1963, R. J. Acheson, P. banksiana , 13, 4♀ (CNC); Wiarton, 1.VII.1962, G. Thorpe, 13 (CNC). Québec: Fabre, 5.VII.1963, L. A. Kelton, Picea glauca , Pinus banksiana , 193, 16♀ (CNC); Kazabazua, 15.VIII.1961, D. J. E. Brown, P. banksiana , 1♀ (CNC); Laniel, 3–4, 10–11.VII.1965, L. A. Kelton, W. Gagne, Picea sp. , Pinus banksiana , 383, 63♀ (AMNH, CNC); Missisquoi Co.: Mount Pinacle, nr Frelighsburg, 400 m, 20.VI.1991, M. D. Schwartz, Abies balsamea , 33, 178♀ (CNC); Charlevoix­Est Co.: Port­au­Persil, 1.VIII.1986, Larochelle, Larivière, 2♀ (CNC); Rolled, 15.VIII.1968, W. Gagne, P. banksiana , 13, 6♀ (CNC); Taboret Station, 10.VIII.1963, L. A. Kelton, Pinus sp. , 33, 5♀ (CNC). Saskatchewan: Christopher Lake, 13.VII.1959, P. banksiana , A., J. Brooks, 23, 9♀ (CNC); Hudson Bay, 25.VIII.1954, Brooks, Wallis, 1♀ (CNC); Meadow Lake, 6.IX.1970, L. A. Kelton, P. banksiana , 1♀ (CNC). Prince Albert: 5–7 km N of, on Rte 2, 12.VII.1990, M. D. Schwartz, P. banksiana , 33, 2♀ (CNC); 23.VII.1959, A., J. Brooks, 7♀ (CNC). Torch River: Campground, E of Rte 106, 12.VII.1990, M. D. Schwartz, P. banksiana , 93, 17♀ (CNC); 11.VII, 12.VIII.1950, L. A. Konotopetz, P. banksiana , 33, 1♀ (CNC). White Fox, 24.VII.1950, L. A. Konotopetz, P. banksiana , 2♀ (CNC). Yukon Territory: Carcross: 28.VI.1982, G. G. E. Scudder, listed in Scudder 1977, 2♀ (UBC); 1, 9.VIII.1983, L. A. Kelton, P. contorta , 13, 14♀ (CNC). Moose Creek, 18.VII.1982, 28.VII.1983, L. A. Kelton, Pinus sp. , 33, 21♀ (CNC); Morley River, L. A. Kelton, P. contorta , 2♀ (CNC); Squanga Lake, L. A. Kelton, P. contorta , 2♀ (CNC); Tagish, 11.VIII.1983, L. A. Kelton, 13 (CNC); Tatchun, 11.VIII.1983, L. A. Kelton, P. contorta , 13 (CNC); Watson Lake, 13.VIII.1982, L. A. Kelton, 1♀ (CNC); Whitehorse, 30.VII.1982, L. A. Kelton, P. contorta , 63, 9♀ (CNC). USA: Arizona: Coconino Co.: Kaibab Lake, Kaibab National Forest, 16.VIII.1967, L. A. Kelton, 2♀ (CNC). San Francisco Mountains: Coconino National Forest, 9650 ft, 14, 15.VII.1968, L. A. Kelton, 13, 1♀ (CNC); 9.VIII.1929, E. D. Ball, Lepidopsallus longirostris paratypes, 43, 4♀ (USNM). California: El Dorado Co.: Lake Tahoe, 8.VIII.1937, Drake, Andre, Lepidopsallus hesperus paratypes, 23 (USNM); Strawberry, 8.VIII.1929, L. D. Anderson, 13 (KU). Inyo Co. : 20 mi W of Bishop, 9500 ft, 2.VIII.1954, J. D. Lattin, 33 (OSU); Round Valley, 27.VII.1947, R. H. Beamer, 2♀ (KU). Lassen Co.: Bogard Campground, 24 mi W of jct. Hwys 36 and 44, 10.VIII.1980, G. M. Stonedahl, P. ponderosa , 1♀ (AMNH). Mono Co.: Inyo National Forest, D. Poole, 2♀ (LACM). Nevada Co.: Sagehen Creek Station nr Hobart Mills, 20.VIII.1969, W. Gagne, 2♀ (UCB); Truckee, Tahoe National Forest, 29.VIII.1967, L. A. Kelton, Artemisia sp. , 1♀ (CNC). Siskiyou Co.: Medicine Lake, on Medicine Lake Rd, G. M. Stonedahl, J. D. McIver, P. contorta : 1.4 mi S of, 19.VII.1985, 23, 1♀ (AMNH); 2.5 mi N of, 18.VII.1985 13 (AMNH). 6.6 mi S of Lava Beds National Monument, Lava Beds–Medicine Lake Road, 18.VII.1985, G. M. Stonedahl, J. D. McIver, P. contorta , 43, 7♀ (AMNH). Tuolumne Co.: Yosemite National Park, McCabe Lakes, 2.VIII.1939, R. L. Usinger, 1♀ (UCB). Colorado: Boulder Co.: Nederland, Roosevelt National Forest, 4.VIII.1968, L. A. Kelton, P. albicaulis , 173, 14♀ (CNC). Gilpin Co.: Pinecliffe, 12.VIII.1973, J. C. Schaffner, 1♀ (TAM). Grand Co.: 3.4 mi W of Fraser, St. Louis Creek Campground, 9000 ft, 16.VIII.1980, G. M. Stonedahl, P. contorta , 23 (AMNH). Jackson Co.: Crowley, nr, at Jackson Co. and Wyoming State line on Rte 125, 4.VIII.1979, J. T. Polhemus, 4♀ (JTP). Larimer Co.: Fish Creek Picnic ground, Pingree Park Road, 46 mi W of Fort Collins. 7700 ft, 14.VII.1986, R. T. Schuh, J. T. Polhemus, 13, 1♀ (AMNH). Las Animas Co.: 1 mi S of Cucharas Pass, 9000 ft, 29.VIII.1982, D. A., J. T. Polhemus, 1♀ (JTP). Park Co.: Wilkerson Pass, Pike National Forest, 28.VIII.1968, P. albicaulis , 13 (CNC). Connecticut: New Haven Co.: North Haven, 2.VII.1924, J. R. Torre­Bueno, 13 (USNM). Idaho: Benewah Co.: 4 mi E of Emida on Charlie Creek Road, 8.VII.1979, G. M. Stonedahl, 13 (AMNH); 4 mi W of Emida on Hwy 6, 9.VII.1979, G. M. Stonedahl, P. contorta , 13 (AMNH). Vicinity Emida, St. Joe National Forest, T42N R53W, 3000 ft, 5.VIII.1986, Schuh, Schwartz, Stonedahl, 2♀ (AMNH). Bonner Co.: Pend Orielle River, Laclede, 4.VII.1966, W. Gagne, J. Haddock, P. flexilis , 23 (UCB). Fremont Co.: Big Springs, 6.VIII.1931, H. H. Knight, Pinus sp. , Lepidopsallus hesperus paratypes, 13, 4♀ (CNC, USNM). Teton Co.: Pine Creek Pass, 6.VIII.1972, L. A. Kelton, P. contorta , 2♀ (CNC). Iowa: Clay Co.: Peterson, 16.VI.1981, D. A., J. T. Polhemus, 13, 2♀ (JTP). Story Co.: Ames: 23.VI.1933, H. M. Harris, 13 (TAMU); H. H. Knight: 20.VI.1932, 13, 4♀ (USNM); 23.VI.1927, including Atractotomus crataegi paratypes, 353, 6♀ (CNC, TAMU, USNM); 6.VII.1927, 33, 14♀ (USNM); 11.VII.1928, 1♀ (USNM); 30.VI, 5.VII.1951 J. Laffoon, Pinus sp. , 4♀ (AMNH); 19, 20, 27.VI.1962, J. C. Schaffner, Pinus strobus , P. sylvestris , 4♀ (TAMU); 23.VI.1950, J. A. Slater, 13 (AMNH). Maine: Aroostock Co.: Smyrna Mills, 26.VI.1952, 13 (USNM); Portage, 27.VI.1950, 1♀ (USNM); Wade, 30.VI.1950, Washburn, 1♀ (USNM). Hancock Co.: Bar Harbor, 3.VIII.1943, A. E. Brower, 1♀ (USNM). Maryland: Saint George’s Co.: Branchville to Beltsville, 4.VI.1914, W. L. Mc­ Atee, 1♀ (USNM). Massachusetts: Barnstable Co.: Dennis, 4.VII.1954, J. W. Green, 13 (CAS). Essex Co.: Pigeon Cove, 28.VII.1916, C. E. Olsen, holotype Lepidopsallus olseni , 1♀ (USNM). Michigan: Marquette Co.: Marquette, 28.VIII.1988, 13 (USNM). Schoolcroft Co.: Grand Marais, 28.5 km S of, 17.VIII.1993, M. D. Schwartz, P. banksiana , 23, 3♀ (CNC); Seney, 7 km W of, just N of Rte 28, 17.VIII.1993, M. D. Schwartz, P. banksiana , 2♀ (CNC). Minnesota: Brown Co.: New Ulm, 20.VI.1921, H. H. Knight, 1♀ (USNM). Mississippi: Holmes Co.: Tchula, 18.V.1931, H. G. Johnston, 43 (TAMU). Montana: Deer Lodge Co.: Georgetown Lake on Rte 10A, 6500 ft, 9.VIII.1986, Schuh, Schwartz, Stonedahl: P. contorta , 73, 15♀ (AMNH); Potentilla fruticosa (Rosaceae) , 23, 1♀ (AMNH). Gallatin Co.: Moose Flat Campground, 26 mi S Bozeman Hot Springs, Rte 191, 5700 ft, 10.VIII.1986, Schuh, Schwartz, Stonedahl, P. contorta , 23, 3♀ (AMNH). Glacier Co.: Glacier National Park, 10 mi W of Babb, 8.VIII.1969, P. W. Oman 1♀ (OSU). Granite Co.: 15 mi S of Drummond on Rte 10A, 5000 ft,. VIII. 9, 1986, Schuh, Schwartz, Stonedahl, P. contorta , 13 (AMNH). Park Co.: Rte 212 at WY Border, 7750 ft, 11.VIII.1986, Schuh, Schwartz, Stonedahl, P. contorta , 1♀ (AMNH). Nevada: Washoe Co.: Little Valley Research Area, 4.5 mi SW of Washoe, T16N R19E Sec2 NE 1/4, 6200 ft, 4.VIII.1982, M. D. Schwartz, P. contorta , 213, 19♀ (AMNH). New Hampshire: Coos Co.: Gorham, 8.IV.1978, L. A. Kelton, 1♀ (CNC). New York: Essex Co.: Lake Placid, M. D. Schwartz, P. sylvestris : 4.VII.1991, 103, 35♀ (CNC); Patch Lane, 595 m, 3.VII.1991, 53, 11♀ (CNC). Whiteface Mountain, 1493 m, 4.VII.1991, M. D. Schwartz, Abies balsamea , 53, 5♀ (CNC). Nassau Co.: Roslyn, Fine Arts Museum and Gardens on Rte 25A, 10.VI.1987, 20.VI.1988, 27.VI.1986, M. D. Schwartz, P. sylvestris , 33, 8♀ (AMNH, CNC). Suffolk Co.: East Quoque, Ice Pond Road, 18.VII.1988, M. D. Schwartz, Pinus rigida , 13, 4♀ (AMNH, CNC). Tompkins Co.: Ithaca, Cornell University Campus, 1.VII.1978, A. G. Wheeler, Jr., P. echinata , P. resinosa , 13, 3♀ (PDA). Ulster Co.: Rte 28A near outlet of Ashokan Reservoir, 13.VII.1979, P. Wygodzinsky, K. Schmidt, Pinus sp. , 33, 20♀ (AMNH). Oregon: Baker Co.: Boulder Park and Cougar Meadow, 20 mi E of Medical Springs, Wallowa Mountains, 4990 ft, 3.VIII.1986, Schuh, Schwartz, Stonedahl, P. ponderosa , 13, 14♀ (AMNH). Two Color Campground, 17 mi E of Medical Springs, Wallowa Mts., 4650 ft, 2.VIII.1986, Schuh, Schwartz, Stonedahl, P. contorta , 23, 4♀ (AMNH). West Eagle Meadow, 20 mi E of Medical Springs, Wallowa Mts., 4700 ft, 3.VIII.1986, Schuh, Schwartz, Stonedahl, P. contorta , 33, 4♀ (AMNH). Coos Co.: 7.3 mi E of Bandon, 15m, 12.VII.1979, R. T., Joe Schuh, 1♀. Curry Co.: 2 mi N of Port Oxford, 30 m, 12.VII.1979, R. T., Joe Schuh, P. contorta , 133, 25♀ (AMNH). Brookings, 4 mi N of, 60 m, 12.VII.1979, R. T., Joe Schuh, 1♀ (AMNH). Deschutes Co.: Three Creeks Meadow, 1.IX.1977, Lattin, P. contorta , 2♀ (OSU). Grant Co. : Funny Bug Basin, T17S R32E Sec 19, Malhuer National Forest, 23.VII.1979, M. D. Schwartz, Juniperus occidentalis (Cupressaceae) , 1♀ (AMNH). Hood River Co.: Trillium Lake, 3500 ft, 12.IX.1978, J. D. Lattin, P. contorta , 13 (OSU). Klamath Co.: Cleetwood Cove Trail, Crater Lake National Park, 3.VIII.1979, M. D. Schwartz, P. contorta , 13, 2♀ (AMNH); Crescent, 1 mi W of, 4500 ft, J. D. Lattin, M. D. Schwartz, P. contorta , 53, 4♀ (AMNH, OSU); Meadow Spring, Winema National Forest, T27S R9E S16, 5500 ft, 17.VII.1979, M. D. Schwartz, P. contorta , 13 (AMNH). Lincoln Co.: Beverly Beach State Park, 5.VII.1979, G. Stonedahl, P. contorta , 13, 1♀ (AMNH); Fogarty Creek State Park, 5.VII.1979, G. Stonedahl, M. D. Schwartz, P. contorta , 283, 18♀ (AMNH); near Yaquina Light House, 5.VII.1979, M. D. Schwartz, G. M. Stonedahl, P. contorta , 23, 3♀ (AMNH, OSU); South Beach State Park, in campground and trail to beach, 29.VI.1994, M. D. Schwartz, A. E.­Martinez, P contorta , 153, 12♀ (CNC). Wallowa Co.: Miram Lake Trail, Wallowa­Whitman National Forest, 5590–7370 ft, 18.VIII.1979, M. D. Schwartz, P. contorta , 1♀ (AMNH). Pennsylvania: Dauphin Co.: Hershey, Hershey Hotel Grounds, 9.VI.1974, A. G. Wheeler, P. virginiana , 13 (AMNH). Fishing Creek Valley School, Rte 443, 9.VI.1976, A. G. Wheeler, P. virginiana , 13 (AMNH). Monroe Co.: Greenview Road, 3 mi E of Brodheadsville, 6.VII.1985, Schuh, Schwartz, Stonedahl, P. resinosa , 3♀ (AMNH). Tennessee: Cocke Co.: Rtes 25, 70 between 107N and 107S, 20.V.1979, Wygodzinsky, Schuh, Schmidt, 13, 3♀ (AMNH). Grainger Co.: Rte 25E, N of Cherokee Lake, 20.V.1979, Wygodzinsky, Schuh, Schmidt, 13 (AMNH). Knox Co.: Knoxville, University of Tennessee Campus, 27.V.1985, A. G. Wheeler, Jr., Picea abies , 13 (PDA). Virginia: Washington Co.: Bristol, 26.V.1985, T. J. Henry, A. G. Wheel­ er, Pinus virginiana , 23, 2♀ (USNM). Wisconsin: Oneida Co. : American Legion State Forest, T39N, R8E, 17.VII.1957, P. A. Jones, P. banksiana , 1♀ (USNM). Wood Co.: Griffith Street Nursery, 19.VI.1951, R. D. Shonefelt, 1♀ (AMNH). Wyoming: Park Co.: Shoshone National Forest, 15.VIII.1927, H. H. Knight, Lepidopsallus hesperus paratype, 1♀ (USNM). Shoshone Co.: Fox Creek Campground, 6.9 mi E of Cooke City on Rte 212, 7250 ft, 11.VIII.1986, Schuh, Schwartz, Stonedahl, P. contorta , 1♀ (AMNH). Teton Co.: Colter Bay Village, 2 mi SE of, on Rte 89, Grand Teton National Park, 7000 ft, 22.VII.1981, M. D. Schwartz, P. contorta , 13, 1♀ (AMNH). Grand Teton National Park, 6900 ft, 18.IX.1980, J. D. Lattin, P. contorta , 1♀ (OSU).

Phoenicocoris strobicola (Knight)

Figures 1N, 3 , 19 , 24

Psallus strobicola Knight, 1923: 467 (n.sp.); Wheeler, et al., 1983: 138 (list, host).

Phoenicocoris strobicola: Kerzhner, 1962: 381 (n.comb.); Schuh, 1995: 415 (catalog).

DIAGNOSIS: Similar to the sexually dimorphic obscurellus in the relatively elongate body of the male, the narrow, scalelike setae on the dorsum and lateral margins of body, and the long antennal segment 2, but distinguished by the much paler antennae and legs, and the gonopore sclerite subequal to the length of the secondary gonopore and the distal margin of the gonopore with prominent spinules (fig. 24).

REDESCRIPTION: GENERAL ASPECT: elongate (male) to moderately elongate (female); total length male 3.28 (3.13–3.50), female 2.95 (2.80–3.15); length to cuneal fracture male 2.28 (2.13–2.34), female 2.13 (1.98–2.31); coloration dark brown to black; antennae, legs, except coxae and tarsi pale yellow; dorsum, including vertex and pronotum, with densely distributed, broad, apically pointed, silvery scalelike setae and densely distributed, suberect to reclining, brown simple setae; thoracic pleura with scalelike setae. HEAD: width male 0.65 (0.62–0.69), female 0.64 (0.61–0.65); vertex width male 0.28 (0.27–0.29), female 0.30 (0.29–0.31); region anterior to antennal insertion elongate, anteocular length male 0.20 (0.19–0.21), female 0.21 (0.20–0.22); eyes large, 89% of head height, ventral margin of antennal insertion level dorsal to ventral margin of eye; antennal measurements male 0.19 (0.18–0.21): 0.92 (0.84–0.98): 0.57 (0.53– 0.64): 0.35 (0.33–0.36), female 0.18 (0.17– 0.19): 0.84 (0.79–0.89): 0.52 (0.50–0.55): 0.34 (0.32–0.36); segment 2 is 28% longer than head width across eyes; labium yellow, reaching sternite 6 in both sexes; labial length male 1.36 (1.31–1.41), female 1.36 (1.26–1.42); length of segment 4 male 0.36 (0.35–0.38), female 0.35 (0.33–0.38). THO­ RAX: width male 0.96 (0.93–1.05), female 0.94 (0.90–0.98); length male 0.47 (0.44– 0.50), female 0.45 (0.43–0.48); mesoscutum broadly exposed. HEMELYTRA: maximum width male 1.24 (1.13–1.39), female 1.23 (1.18–1.31); lateral margins subparallel.

BIOLOGY: Eastern white pine ( Pinus strobus Linnaeus ) ( Pinaceae ) is the breeding host. The occurrence on other plants are sitting records as adults disperse to nearby plants when populations are high. Based on field work (T. J. Henry, personal commun.) P. strobicola will not be found at a particular locality unless white pines are present.

DISTRIBUTION: An eastern Nearctic species found from Nova Scotia west to Ontario and Minnesota in the north, south to West Virginia and Illinois. Wheeler and Henry (1988) also listed Ohio and Oklahoma within the distribution this species.

DISCUSSION: The discovery of a female we collected on Abies grandis (Dougl.) Lindl. in the Wallowa Mountains of eastern Oregon apparently represents a contaminate in collecting vials or a mislabeled specimen considering the large range disjunction and host plant association of this one specimen. Regardless, the long, spatulate parempodia, long labium, silvery scalelike setae on the dorsum and lateral aspect, and pale yellow appendages substantiate our identification.

SPECIMENS EXAMINED: CANADA: New Brunswick: Fredericton, 28.VI.1976, 1.VIII.1966, L. A. Kelton, Solidago sp. , 13, 2♀ (CNC); Kennebec Banks, 23.VI.1976, L. A. Kelton, Pinus strobus , 123, 5♀ (CNC); Kouchibouguac National Park, 23.VII.1977, D. J. E. Brown, Pinus sp. , 23, 1♀ (CNC); Waterford, 1.VIII.1966, L. A. Kelton, P. strobus , 43, 13♀ (CNC). Nova Scotia: Halifax, 22.VII.1975, L. A. Kelton, Picea sp. , 1♀ (CNC); Ingonish, 2.VIII.1976, L. A. Kelton, Pinus sp. , 13 (CNC). Kentville, 3–6.VII.1976, 15–17.VII.1968, L. A. Kelton, Pinus sp. , 103, 2♀ (CNC); Mount Unlacke, 13–14.VII.1966, L. A. Kelton, 23 (CNC); Woodville, 20.VII.1966, L. A. Kelton, Larix sp. , 3♀ (CNC). Ontario: Almonte, 28.VI.1979, D. J. E. Brown, 33, 2♀ (CNC); Buckhorn, 28.VI.1968, G. Thorpe, P. strobus , 53, 15♀ (CNC); Footes Bay, 25.VII.1962, Kelton, Thorpe, 2♀ (CNC); Hasting Co., 19.VI.1955, J. F. Brimley, 43, 5♀ (CNC); Huntley, 14.VII.1961, D. J. E. Brown, 33, 1♀ (CNC); Ignace, 5 km W of, on Hwy 17 roadside, M. D. Schwartz, P. banksiana , 13 (CNC); Lakefield, 28.VI.1962, H. Blanchard, 1♀ (CNC); Marmora, 5.VII.1952, J. R. Vockeroth, Pinus sp. , 2♀ (CNC). Nepean, Piney forest, Lafontaine house, M. D. Schwartz, MV light: 18.VII.1991, 143, 4♀; 27.VII.1991, 1103, 33♀ (AMNH, CNC). Nestor Falls, 7–8.VIII.1960, Kelton, Whitney, Pinus sp. , 1♀ (CNC); Norway Point, Lake of Bays, 28.VI.1922, J. McDunnough, Pinus sp. , 53, 4♀ (CNC); Ompah, 28.VI.1979; D. J. E. Brown, 23 (CNC); One Sided Lake, 4– 5.VIII.1960, Kelton, Whitney, 1♀ (CNC). Ottawa, Central Experimental Farm, M. D. Schwartz: 19.VI.1991, Picea glauca , Pinus sylvestris , 73, 6♀ (CNC); ca Saunders Building, 17.VI.1999, 73, 1♀ (CNC). Parry Sound, 26.VII.1962, Kelton, Thorpe, 1♀ (CNC). Saint Lawrence Islands National Park: Gordon Island, 7.VIII.1976, W. Reid, 1♀ (CNC); Grenadier Island Centre, 27.VII.1975, E. Wilson, 13 (CNC); Thwartway Island, 19.VII.1976, W. Reid, 1♀ (CNC). Stirling, 26.VI.1962, G. Thorpe, Pinus sp. , 53, 8♀ (CNC); Stittsville, 12.VII.1979, D. J. E. Brown, 13 (CNC). Québec: Fabre, 5.VII.1961, W. Gagne, Picea gluaca , 93, 1♀ (CNC); Ladysmith, 24.VII.1958, L. A. Kelton, 13 (CNC); Laniel, 3– 4.VII.1963, L. A. Kelton, 13 (CNC); Shawville, 22.VII.1958, L. A. Kelton, Pinus strobus , 123 (CNC). USA: Illinois: Jo Daviess Co.: Galena, 30.VI.1922, Dozier, Mohr, P. strobus , 1♀ (AMNH). Maryland: Prince George’s Co.: College Park, University of Maryland, 4.VI.1988, T. J. Henry, A. G. Wheeler, Jr., Pyracantha sp. , 1♀ (USNM). Massachusetts: Berkshire Co.: Lake Buell, Hurstville, 2.VII.1930, J. R. Torre­Bueno, 2♀ (KU). Michigan: Emmet Co.: 1 mi S of Pellston, 30.VII.1957, R. E. Beer, 2♀ (KU). Minnesota: Carlton Co.: Elkhorn Creek, 18.VIII.1920, H. H. Knight, 13 (USNM). Ramsey Co.: St. Anthony Park, 22.VIII.1924, H. H. Knight, 13 (USNM). New York: Albany Co.: Rensselaerville, Huyck Preserve, 29.VĪ 2.VII.1977, R. T. Schuh, Pinus strobus , 33, 5♀ (AMNH). Essex Co.: Lake Placid, 4.VII.1991, M. D. Schwartz, Picea glauca , 1♀ (CNC). Nassau Co.: East Meadow near Rte 25 on Bluebird Drive, M. D. Schwartz, 6, 10, 16.VI, 16.VIII.1986, Picea abies , Pinus strobus , Tsuga canadensis , 343, 13♀ (AMNH); Flower Hill near Rte 25A on Ridge Drive East, 10.VI.1986, 8.VII.1985, M. D. Schwartz, P. strobus , 1♀, Picea glauca , 1♀ (AMNH); Roslyn, Cemetery on Rte 25A, 23.VI.1986, M. D. Schwartz, Pinus strobus , 53, 14♀ (AMNH); Roslyn, Fine Arts Museum and Gardens on Rte 25A, 10.VI.1986, 1987, 18, 20.VI.1988, M. D. Schwartz, P. sylvestris , Betula sp. (Betulaceae) , P. strobus , 43, 7♀ (AMNH). Rockland Co.: South Nyack, Ross Avenue, 18.VI.1988, M. D. Schwartz, Pinus sp. , 13 (AMNH). Suffolk Co.: Bayshore, Brentwood Road, 22, 23.VI.1985, M. D. Schwartz, P. strobus , 143, 13♀ (AMNH). Tompkins Co.: Ithaca: H. H. Knight, 29.VI.1920, P. strobus , Psallus strobicola paratype, 43, 2♀ (CAS, CNC, USNM); A. G. Wheeler, Jr.: Cornell University Campus, 1.VII.1978, Pinus mugo , 2♀ (PDA); 0.5 mi NW of, 30.VI.1978, P. strobus , 13 (PDA). Ringwood Road, 13.VII.1920, 1♀ (USNM). Taughannock Falls, 2.VII.1920, 1♀ (USNM). North Carolina: Henderson Co.: Hendersonville, 1.VI.1907, F. Sherman, 13 (USNM). Macon Co.: Highlands, 35°3.2'N 83°11.3'W, 3850 ft, 5, 18.VII.1958, J. L. Laffoon, light, 33 (USNM). Union Co.: Monroe, 28.V.1973, A. G. Wheeler, Jr., P. strobus , 13 (PDA).? Oregon: Baker Co.: Wallowa Mountains, Boulder Campground and Cougar Meadow, 20 mi E of Medical Springs, 4490 ft, 3.VIII.1986, Schuh, Schwartz, Stonedahl, Abies grandis , 1♀ (AMNH). Pennsylvania: Blair Co.: Altoona, Pleasant Valley Nursery, 11.VI.1972, S. M. George, Picea abies , 13 (PDA). Centre Co.: 2 mi N of State College, 10.VI.1977, Schuh, Henry, Wheeler, 3♀ (AMNH). Cumberland Co.: Camp Hill, 12.VI.1973, A. G. Wheeler, Jr., Pinus sylvestris , 1♀ (PDA). Dauphin Co.: Harrisburg, Agriculture Building, 10, 30.VI, 1973, 1974, A. G. Wheeler, Jr., P. strobus , 13, 1♀ (PDA); Hershey Hotel grounds, 12, 27.VI.1973, T. J. Henry, B. R. Stinner, A. G. Wheeler, Jr., P. sylvestris , P. strobus , 23, 7♀ (PDA). Erie Co.: Erie County Hospital, 4.VII.1972, F. Negley, H. Wolff, P. strobus , 43 (PDA). Franklin Co.: Mainsville, Snipe’s Nursery, 16.VI.1972, Hauser, P. strobus , 1♀ (PDA). Huntingdon Co.: Greenwood Furnace, 28.VI.1990, A. G. Wheeler, Jr., P. strobus , 43, 1♀ (PDA). Monroe Co.: Long Pond, nr, 600 m, 41°01'N 75°28'W, M. D. Schwartz, P. rigida , 1♀ (CNC); Stroudsburg, La Bar’s Nursery, 26.VI.1973, A. G. Wheeler, Jr., P. strobus , 23 (PDA). Montgomery Co.: Barnes Arboretum, Merion Station, 2.VI.1982, A. G. Wheeler, Jr., P. strobus ‘nana’, 23 (PDA); Forest Hills Cemetery, Philadelphia, 7.VI.1973, J. F. Stimmel, Picea abies , 13 (USNM); Manufacturers Golf and Country Club, 13.VI.1973, A. G. Wheeler, Jr., Pinus strobus , 3♀ (PDA). Washington Co.: Iannett’s Nursery, 21.VI.1973, T. J. Henry, P. strobus , 13, 2♀ (PDA). Wayne Co.: Curtis Nursery, nr Callicoon, 3.VIII.1976, 1♀ (PDA). Rhode Island: Providence Co.: Providence, 1♀ (USNM). Tennessee: Knox Co.: Knoxville, University of Tennessee Campus, 27.V.1985, T. J. Henry, A. G. Wheeler, Jr., P. strobus , 43, 2♀ (PDA, USNM). Virginia: Albemarle Co.: Charlottesville, University of Virginia Campus, 23.V.1986, A. G. Wheeler, Jr., P. strobus , 3♀ (PDA). Augusta Co. : I­81, Verona exit, nr Staunton, 29.V.1973, A. G. Wheeler, Jr., P. strobus , 5th instar, 13 (PDA). Rockingham Co.: Harrisonburg, Rte 33, 25.V.1985, T. J. Henry, A. G. Wheeler, P. strobus , 23 (USNM). Washington Co.: Bristol, 26.V.1985, T. J. Henry, A. G. Wheeler, P. strobus , 33 (USNM). West Virginia: Greenbrier Co.: White Sulphur Springs, 25.V.1978, A. G. Wheeler, Jr., P. strobus , 3♀ (PDA).

Figures

Fig. 3. Dorsal habitus photographs. See appendix 1 for localities.

Fig. 16. Scanning electron micrographs of Phoenicocoris pallidicornis, n.sp., Falcon Lake, Manitoba. A. Head and thorax, lateral view. B. Scalelike setae, dorsal view. C. Hind femora, dorsobasal view. D. Pretarsus, apical view. Scales as indicated.

Fig. 24. Male genitalia of Phoenicocoris species—pallidicornis, Manitoba: Falcon Lake; ponderosae, Colorado: 3 mi S of Guffey; rostratus 1, New York: E. Qogue; rostratus 2, Iowa: Ames; strobicola, New Brunswick: Kennebec. Scale = 0.10 mm.

Fig. 17. Scanning electron micrographs of Phoenicocoris ponderosae, n.sp., Rustler Park, Arizona. A. Scalelike setae, dorsal view. B. Hind femora, dorsodistal view. C. Pretarsus, ventral view. Scales as indicated.

Fig. 18. Scanning electron micrographs of Phoenicocoris rostratus (Knight), Fish Creek, Colorado. A. Head and thorax, lateral view. B. Scalelike setae, dorsal view. C. Pretarsus, apical view. D. Hind femora, dorsodistal view. Scales and magnifications as indicated.

Fig. 19. Scanning electron micrographs of Phoenicocoris strobicola (Knight), Nepean, Ontario. A. Head and thorax, lateral view. B. Scalelike setae, dorsal view. C. Hind femora, dorsobasal view. D. Hind femora, dorsodistal view. E. Pretarsus, apical view. F. Vesica, distal view. Scales as indicated.