Engistanoxia dessertineae Lacroix, 2002

Sehnal, Richard, 2014, Engistanoxia sagala sp. nov. from Kenya (Coleoptera: Scarabaeidae: Melolonthinae: Melolonthini), Acta Entomologica Musei Nationalis Pragae 54 (2), pp. 563-569 : 568-569

Engistanoxia dessertineae Lacroix, 2002

( Figs 8–14 )

Engistanoxia dessertinae Lacroix, 2002: 159 (primary description); LACROIX (2010): 198 (redescription, catalogue).

Type locality. Kenya, Makueni county, Kibwezi.

Type material examined. HOLOTYPE: J ދ KENYA XII-93 / Kibwezi / Werner leg. [p] // HOLOTYPE [p, red label] // Engistanoxia / dessertinei n.sp [hw] / M. LACROIX det. 2002 [p] // Engistanoxia / dessertineae Lacroix, 2002 / M. Lacroix det. [p] // Engistanoxia / dessertineae [hw]ތ ( MNHN).

Redescription. Male (holotype). Body length 25.0 mm (without pygidium). Body elongate, gradually widening posteriorly. Dorsal surface and abdomen reddish brown, legs partly darker to blackish brown, antennae pale brown, elytra pruinose ( Fig. 8 ).

Head. Clypeus 2.3 times wider than long, on lateral margins strongly round and with anterior margin medially emarginate ( Fig. 9 ). Spaces between scales on clypeus and frons amount at most to half width of macrosetae. Eye canthus long, with macrosetae in two irregular rows; anterior columnar macrosetae long, posterior columnar macrosetae half as short and half as many as anterior scales. Antennae decamerous. Antennal club hexamerous, anteriorly gently cambered, 1.7 times longer than antennal shaft (antennomeres 1–4 combined). Antennomere I cambered, at apex evenly rounded and with long setae, antennomere IV prolonged basally in lamela, reaching the ¿rst half of antennomere V. Apex of antennal club with evenly distributed ¿ne hairs ( Fig. 10 ). Terminal maxillary palpomere widest in two thirds, apically truncate and with a terminal tubercle on inner, more angular side, on dorsal surface with a large oval alutaceous area widening in two thirds of length ( Fig. 11 ).

Pronotum weekly convex, approximately nonagonal ( Fig. 12 ), 1.4 times wider than long, widest in front third, with distinct impressed medial line. Anterior and posterior angles round but discernible. Lateral margins of pronotum slim, strongly curved up, in sublateral view appearing smooth but crenulate, with a yellow hook-shaped macroseta issuing from each crenulation ( Fig. 15 ). Surface densely punctate; punctures unevenly distributed, spaces between them smooth and matte; each puncture bearing pointed scale as long as scales on clypeus and frons. Scales forming groups oriented mostly toward midline.

Scutellar shield on periphery with round punctures which bear yellowish-white scales of same size; in basal corners group of densely packed round and prolonged macrosetae and ¿ne hairs; center scaly, on sides with facets devoid of punctures and scales.

Elytra convex, elytron 2.6 times longer than wide. Crests of costae glabrous, their sides and intervals unevenly covered by punctures and scales similar to those on pronotum. Spaces between punctures and scales ¿nely rugate and matte. Lateral margins with hairs twice as long as those on pronotal margins.

Macropterous.

Legs. All femora shiny, irregularly punctate, macrosetae long. Tibiae covered by conspicuous wide scales spaced at most at scale width. Inner tooth of pretarsus half the length of apical tooth. Meso- and metatibiae deeply punctate, macrosetae ¿ne, long and errect.

Abdominal ventrites dark reddish brown, matte, ¿nely punctate, covered by oval yellow scales and a row of semierect yellowish macrosetae. Pygidium slightly concave, shallowly punctate, with yellowish-white caudally oriented scales posteriorly grading into ¿ne, long, compact yellowish-white macrosetae.

Male genitalia ( Figs 13–14 ). Aedeagus symmetrical. Phallobase in lateral view with three extensions, ventrolaterally slightly excised. Parameres forming a symmetrical lobe with an open medial oval summit and an oval concavity in its center. Dorsal part of lobe steeply elevated and then reclined at angle ventral to base.

Figures

Figs 8–15. Engistanoxia dessertineae Lacroix, 2002, holotype male. 8 – habitus, dorsal view; 9 – head, dorsal view; 10 – antenna, dorsal view; 11 – maxillary palp, dorsolateral view; 12 – pronotum, dorsal view; 13 – aedeagus, dorsal view; 14 – aedeagus, lateral view, 15 – lateral margin of pronotum, dorsal view. Scale 1 mm.