Ummidia carabivora (Atkinson, 1886)

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia carabivora (Atkinson, 1886) Figs 10 , 11 , Map 3

Pachylomerus carabivorus Petrunkevitch, 1929: 511; HOLOTYPE: 1 ♀ (A-47) from Chapel Hill, North Carolina, United States, 35.9131 -79.0557 6, 147 m a.s.l., deposited in USNM Marx Collection, examined.

Pachylomerus carabivorus emarginatus Atkinson, 1886: 134, syn. nov.

Diagnosis.

Ummidia carabivora can be differentiated from all other species of Ummidia by the pars cephalica which extends 4/5 (7/10 in other species) the length of the carapace and raised dorsally at the fovea which is 1/3 (1/4-1/5 in other species) the width of the carapace. Males can further be distinguished from all other eastern United States species with tarsal combs by tibia I being relatively more spinose both on the prolateral and retrolateral aspects. Males disperse from June to July.

Description of female holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. General coloration. Carapace, chelicerae, and legs very dusky red 10R 2.5/2. Abdomen very dark brown 7.5YR 2.5/3, spinnerets dark brown 7.5YR 3/4. Cephalothorax. Carapace 9.07 long, 8.3 wide. Pars cephalica 7.21 long. Foveal groove procurved and extremely broad, 1.11 long, 2.81 wide. Eye tubercle low. AER very procurved PER very slightly procurved. Eye group 1.11 long, 2.1 wide, AME 0.46, PME 0.31, ALE 0.56, PLE 0.37. Sternum sparsely setose around anterior edges with posterior fringe, STRl 5.41, STRw 5.3. Chelicerae with anterior row comprising six teeth, posterior margin with six teeth. Palpal endites with 25 cuspules spread across proximal half of endite and 37 cuspules distally, ENDw 1.98, ENDl 3.38. Labium with five cuspules, LBw 1.93, LBl 1.36. Rastellum with many strong spines on process. Abdomen. Evenly setose. Legs. F1 5.54; F1w 1.91; P1 3.37; Ti1 3.41; Mt1 2.34; Tr1 1.32; F3 4.73; F3w 2.51; P3 3.02; Ti3 2.74, Sd3 1.77; Mt3 2.15; Tr3 2.11; F4 5.99; F4w 2.38; P4 3.53; Ti4 3.52; Mt4 3.13; Tr4 1.71. Retrolateral face tarsus IV with defined comb. Pedipalps. PF 4.8, PP 2.67, PTi 3.1, PTr 2.68. Spermathecae straight with slight medial tilt, bulbs facing anteromedially.

Variation, females.

Known only from female holotype specimen.

Description of male exemplar.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Left palp, leg I removed, in vial with specimen. General coloration. Carapace and chelicerae dusky red 10R ¾, legs strong brown 7.5YR 4/6. Abdomen dark brown 10YR 3/3. Cephalothorax. Carapace 8.05 long, 7.16 wide. Pars cephalica 6.85 long. Foveal groove broadly procurved, 1.48 long, 2.34 wide. ME on moderate tubercle. AER procurved. PER straight. Eye group 0.93 long, 1.66 wide, AME 0.48, PME 0.24, ALE 0.47, PLE 0.25. Sternum sparsely setose anteriorly with posterior fringe, STRl 3.98, STRw 4.04. Chelicerae with anterior tooth row comprising four teeth, posterior margin with six teeth. Palpal endites with 19 small cuspules across proximal half of endite face, lacking distal endite cuspules, ENDw 1.58, ENDl 2.86. Labium with five cuspules, LBw 1.47, LBl 1.11. Rastellar process narrow and extended medially; covered in small spines. Abdomen setose, setae very short. Legs. F1 5.71; F1w 1.84; P1 2.92; Ti1 3.68; Mt1 2.61; Tr1 1.31; F3 4.57; F3w 2.05; P3 2.56; Ti3 2.48; Sd3 1.49; Mt3 2.52; Tr3 2.1; F4 5.49; F4w 2.1; P4 2.71; Ti4 3.73; Mt4 3.66; Tr4 1.82. Retrolateral face of tarsus IV with defined comb over central half of tarsus. Leg I spination pattern: TSp 15, TSpv 4, TSrd 0, TSr 3, TSrv 31, MtSp 17, MtSr 14, TrSp 12, TrSr 13. Pedipalps. PTl 3.05, PTw 1.16, Bl 2.3. Embolus evenly curved.

Variation, males

(n = 3). CL 6.16-8.85, 7.69 ± 0.8; CW 5.54-8.23, 6.98 ± 0.78; Cap 4.95-6.85, 6.15 ± 0.6; ENDl 0.59-1.07, 0.86 ± 0.14; ENDw 1.34-1.75, 1.58 ± 0.12; STRl 3.17-4.28, 3.81 ± 0.33; STRw 3.05-4.75, 3.95 ± 0.49; LBl 0.77-1.11, 0.94 ± 0.17; LBw 1.08-1.47, 1.28 ± 0.2; F1 4.35-5.79, 5.28 ± 0.47; F1w 1.38-2.03, 1.75 ± 0.19; P1 2.26-3.18, 2.78 ± 0.27; Ti1 2.92-3.68, 3.38 ± 0.23; Mt1 2.02-2.81, 2.48 ± 0.24; Tr1 1.08-1.48, 1.29 ± 0.12; F3 3.37-5.06, 4.33 ± 0.5; F3w 1.44-2.14, 1.88 ± 0.22; P3 1.84-2.56, 2.31 ± 0.24; Ti3 1.88-2.81, 2.39 ± 0.27; Mt3 1.86-2.75, 2.38 ± 0.27; Tr3 1.61-2.25, 1.98 ± 0.19; F4 4.46-6.37, 5.44 ± 0.55; F4w 1.46-2.2, 1.92 ± 0.23; P4 2.09-3.18, 2.66 ± 0.32; Ti4 2.79-3.85, 3.46 ± 0.34; Mt4 3.13-4.08, 3.62 ± 0.27; Tr4 1.82-2.28, 1.98 ± 0.15; TSp 4-17, 9.33 ± 3.93; TSpv 2-6, 4 ± 1.15; TSr 1-3, 2 ± 0.58; TSrv 25-31, 28.33 ± 1.76; PTl 2.62-6.05, 4.02 ± 1.04; PTw 0.98-1.29, 1.14 ± 0.09; BL 1.96-2.3, 2.13 ± 0.1.

Material examined.

United States: North Carolina: Chapel Hill , 35.9131 -79.0557 6, 147 m a.s.l. (A-47, 1♀, USNM); Cumberland Co: Cool Springs, 35.845 -80.7321 5, 272 m a.s.l. (UMM0691, 1♂, BME) ; Virginia: Virginia Beach Co : Virginia Beach City, Seashore State Park , 36.9185 -76.0523 5, 0 m a.s.l. (UMM0073, 21.vi.1989, 1♂, VMNH); (UMM0025, 23.vi-6.vii-2003, 1♂, R Vigneault, VMNH) .

Figures

Map 3. Locality records for eastern United States species.

Figure 10. Ummidia carabivora (Atkinson, 1886) from Chapel Hill, North Carolina A male habitus illustration UMM 073 B female habitus illustration A 47. Scale bars: 4.0 mm.

Figure 11. Ummidia carabivora (Atkinson, 1886) from Chapel Hill, North Carolina A-D male exemplar (UMM 073) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female holotype (A 47). Scale bars: 1.0 mm.