Chariessa pilosa (Forster), 1771

Opitz, Weston, 2017, Classification, natural history, and evolution of the subfamily Peloniinae Opitz (Coleoptera: Cleroidea: Cleridae). Part VIII. Systematics of the checkered beetle genus Chariessa Perty, Insecta Mundi 2017 (574), pp. 1-42 : 16-25

Chariessa pilosa (Forster), 1771

Figures 12, 13 , 28 , 31, 34 , 36-43, 46, 47, 62.

Chariessa pilosa Forster, 1771: 49 . Lectotype. Gender not known. Type locality: North America. Corporaal, 1950: 276. Opitz, 2014: 25. Forster did not specify as to the number of specimens that were involved in his description. Therefore, I invoke Recommendation 73F of the ICZN (1999) and designate a lectotype for this nominal species

Enoplium marginata , variety of pilosa proposed by Say, 1823: 187. New Synonymy.

Pelonium cincta Spinola, 1844: 356 .

Enoplium onusta Say, 1828 : Table 1, Figure 1 .

Diagnosis. The members of this species resemble superficially those of Chariessa floridana and those of C. texana . In C. floridana specimens the dark pronotal lines are much wider apart (compare Fig. 30, 31 ) and in C. texana members the dark coloration of the pronotal disc is reduced to an anteromedial spot. Also, C. pilosa is widely distributed in the USA and Canada ( Fig. 66 ), whereas C. floridana is known only from the Floridian Key Largo, and C. texana has only been taken from Texas and the panhandle of Oklahoma.

Redescription. Size: Length 10.0 mm; width 4.0 mm. Form: As in Fig. 46 . Mouthparts, cranium, antennae, pterothorax, legs, and abdomen dark brown; pronotum red-yellow at sides, disc middle with two broad black lines between which is a narrow reddish line; elytra dark brown, epipleural margin testaceous beneath. Head: Funicular antennomeres progressively shorter and wider towards capitulum, in males capitulum longer than combined length of funicular antennomeres, capitular antennomeres 9 and 10 with collateral branch ( Fig. 12, 13 ), antennomere 11 oblong, anterior margin sinuous; eyes small, frons wider than width of eye (EW/FW 15/25). Thorax: Pronotum ( Fig. 28 ) slightly transverse (PW/PL 73/70), side margins with small tubercle, disc finely punctate; elytral asetiferous punctation small, profusely distributed throughout disc (EL/EW 230/60); protibial anterior margin spinous. Abdomen: Pygidium oblong / scutiform, faintly incised apically; 5 th visible sternite emarginated; aedeagus ( Fig. 62 ) poorly sclerotized ventrally; phallobasic lobes slightly developed, fimbriate, phallobasic rod poorly developed, phallobasic apodeme explanate at extremity; phallus with subapical sclerotization, phallic apex triangular/subacuminate.

Variation. Size: Length 7.0-15.0 mm; width 2.5-5.7 mm. The sutural and epipleural margins may be flavotestaceous, and the red color of the lateral aspects of the pronotum varies in intensity.

Natural history. Many specimens were reared from woody plants. Among these hardwoods we may include the southern prickly ash [Zanthozylum clava-herculis Linnaeus ( Rutaceae )], the balsam poplar [ Populus balsamifera Linnaeus (Salicaceae) ], the shagbark hickory [ Carya ovata (Mill.) K Koch (Juglandaceae) ], the sugar maple [ Acer saccharum Marchall (Sapindaceae) ], the water birch [ Betula occidentalis Hook. (Betulaceae) ], the chinkapin oak [ Quercus muehlenbergii Engelm. (Fagaceae) ], black walnut [ Juglans nigra Linnaeus (Juglandaceae) ], the bur oak [ Quercua macrocarpa Michx. (Fagaceae) ], the Texas cedar elm [ Ulmus crassifolia Nutt. (Ulmaceae) ], the American elm [ Ulmus Americana Linnaeus (Ulmaceae) ], and the honey locus [ Gleditsia triacanthus Linnaeus (Fabaceae) ] while feeding on Agrilus difficilis Gory (Buprestidaea) .

These beetles were also found on the dead trunk of the post oak [ Quercus stellata Wangenh. (Fagaceae) ], the bitternut hickory [ Carya cordiformis (Wangenh.) K. Koch (Juglandaceae) ], on the wood of the Eastern black walnut [ Juglans nigra Linnaeus (Juglandaceae) ], on the wood of the mockernut hickory [ Carya tomentosa Sarg. (Juglandaceae) ], on the wood of the scarlet oak [ Quercus coccinea Muenchh. (Fagaceae) ], on the wood of the North American fir [ Abies balsamea (Linnaeus) Mill. (Pinaceae) ], on the wood of the Eastern redbud [ Cercis Canadensis Linnaeus (Fabaceae) ], on the wood of the southern red oak [ Quercus falcata Michx. (Fagaceae) ], on the wood of the American persimmon [ Diospyrus virginiana Linnaeus (Ebenaceae) ], on the wood of the wingleaf soapberry [ Sapindus saponaria Linnaeus ( Sapindaceae ], on the wood of aspen [ Populus tremuloides Michx. (Selicaceae) ] eating Agrilus Curtis (Buprestidae) , on the wood of the boxelder [ Acer negundo Linnaeus (Apindaceae) ], on the wood of the downy serviceberry [ Amelanchier arborea (F. Michx.) Fernald (Rosaceae) ], on the wood of the eastern black oak [ Quercus velutina Lam. (Fagaceae) ]. These beetles were also captured on ironwood [ Olneya tesota A. Gray (Fabaceae) ], on peach [ Prunus persica (Linnaeus) Batsh (Rosaceae) ], and on logs of pecan [ Carya illinoiensis (Wangenh.) K. Koch. (Juglandaceae) ]. Some specimens were taken in a Malaise trap and others on vegetation at altitudes ranging from 650 to 680 m.

Distribution ( Fig. 66 ). I examined 1559 specimens from: CANAD A: Alberta; Fort MacLeod; 7 mi S Empress, 5-I-1978, reared from birch log, G. Hilchie: Manitoba; Criddle,?-XII-?, collector not noted; Makinak; Onah, 8-VII-1918, J. B. Wallis; Husavik, 6-VII-1910, J. B. Wallis; Aweme, 20-VII-1912, 21-VI- 1992: Ontario; Barry’s Bay, 3-VIII-1968, on poplar eating buprestids, collector not noted; Kent County, Tilbury,?- VI-1980, K. Stephan; Essex County, Wheatly,?- V-1967, K. Stephan Larson’s Camp, One Sided Lake, 19-VII-1962, UV light, J. C. E. Riotta; Ottawa, 10-VII-1912; Cape Chin, 6-VII-46, on maple; Constance Bay, 14-VII-1954; idem, 22-Vi-1960, cut poplar, A. T. Howden; Rondeau Park, South Point Trail, 31-VI-1985, L. Lesage; Hamilton, 28-VI-14-VII-1982, M. Sanborne; idem, collection date and collector not noted; idem, 1-VI-1968, on log of red oak, collector not noted; Rainy River District, 7-VII- 1926, J. F. Brimley; Prince Edward County, 10-VII-1940, J. F. Brimley; Sudbury, 1988; Laemington, 24- VI-1940, W. J. Brown: Quebec; Dosquet, Lotbiniere, 3-VIII-1985, P. Bélanger; Old Chelseae, summit King Mountain, 12-VII- 1071, 1150 feet, A. T. Howden; Rignaud, Vaudreuil, 17-VI-1982, P. Bélanger; Ste- Agathe, Lotbiniére, 3-VIII-1986, Pierre Bélanger; Duparquet, 27-VI-1937, G. Stace Smith; idem, 2-VII- 1942, G. Stace Smith; idem, 5-VII-1941, G. Stace Smith; idem, 2-VII-1941, G. Stace Smith; idem, 23-VII- 1940, on Abies balsamea, G. Stace Smith ; idem, 28-VI-1942, G. Stace Smith; Queen’s Park, Aylmer, 26- VII-1923, C. B. Huchinga; Terrasse-, Vaudreuil, 20-VI-2012, on trunk of dead Carya cordiformis, P. de Tonnancour ; Summit of Mont Rigaud, Clé Vaudreuil, 26-VI-1998, D. M. Wood; Laniel, July 29, 1935, H. S. Fleming: Saskatchewan; Atto’s Lake, Cut Knife, 18-VI-1940, A. R. Brooks: British Colombia; Salmon Arm, emerged from Betula occidentalis , 17-V-1934, O. R. Leech. UNITED STATES OF AMERICA: Alabama; Mobile County, Mobile,?-V-?, collector not noted; Washington County, Calvert,?- VII-1914, collector not noted,; Blount County, Blountsville, 3-VII-1969, T. King; Jefferson County, Birmingham, 26-VI-1964, at light, H. R. Steeves, Jr.; idem, 30-V-1953, on foliage, H. R. Steeves, Jr.; idem, 27-IV-1954, on foliage, H. R. Steeves, Jr.; Jefferson County, Birmingham, 3-VII-1959, T. King; Jefferson County, Mount Brook, 6-VI-1972, T. King; Walker County, nr. Jasper, Devil’s Ladder, 20-VII- 1973, T. King’ Walker County, Jasper, 9-XI-1978, Tim King; Baldwin County, Daphne, 6-VI-1960, B. K. Dozier; Lee County, Auburn, 15-V-1980, persimmon, D. J. Waters; Hale County, vic Shiloh Cpgd, Talledega National Forest, 22-VI-1999, T. C. MacRae; Madison County, Huntsville, 1-VI-1982, beating, L. L. Lampert: Arkansas; Lonoke County, Carlisle, collection date and collected not noted; Hempstead County, Hope, 12-VI-1960, W. F. Barr; idem, collection date and collector not noted; Washington County, Mount Sequoyah, 20-VII-1938, at light, M. W. Sanderson; Marion County,?- VI-1897, collector not noted; Washington County, Lake Wedington, 8-V-1973, L. L. Borown, Jr.; Hempstead County, 11-VI-1954, J. W. Green; Sebastian County, 27-VI-1983, light, P. Skelley; Fulton County, Mammoth Spring, 28-VI-1988, deciduous forest; Fayetteville, 12-20-VI-1973, E. V. Cage; Crawford County, Lake Fort Smith State Park, 1-VI-1990, D. J. Heffern: California; Tuolumae County, Sonora, 14-VI-1985, Jim Cope: Colorado; Boulder County, 15- VIII-2015, Lindgreen Funnel Trap, USDA APHIS; idem, 15-VII-2016; idem, 3-VI-1016; Douglas County, 22-VII-2008, Lindgrin Funnel, Ethanol lure, Colorado Department of Agriculture; Larimer County, Fort Colins, Alison Hall, 13-VI-2014, ex. black walnut; Larimer County, Owl Canyon, 2-VII-1940, collector not noted; Weld County, Fort Lupton, 21-VII-2016, Lindgreen Funnel, Pityogenes lure, T. C. Woods: Connecticut; New Haven County, 10-VI-1911, Champlain; Litchfield County, Cornwall, 22-VI-1922, Chamberlain; idem, 22-VI-23, collector not noted; idem, 24-VI-1923, C.A. Frost; idem, 24-VI-1923, Chamberlain; idem, 1-VI-1922, K. F. Chaimberlain; idem; 8-VII-1928, K. F. Chamberlain; Litchfield County, 11-?-1921, K. F. Chamberlain; Fairfield County, Stanford, collection date and collector not noted: Delaware; Taylor County, 2 mi SE Salem, 20-IV-1976, R. L. Penrose; Sussex County; Bridgeville, 15-VII- 1958: Florida; Marion County, Route 316, Oklawaha River, under Bridge, 5-V-2002, M. H. Rabovsky; 2.5 SSE Micanopy, 2-13-IV-2013, Malaise trap, D. B. Wahl; Indian River County,. 5 mi W I-95, 29-III-2- IV-1976, collector not noted; Leon County, Tallahassee, 6-29-1902, UV trap, G. J. Wibmer & C. W. O’Brien; idem, 4-VII-2002, G. J. Wibmer; Polk County, Lake Marion Estate, 1-V-1999, M. C. Thomas; Polk County, Highway 542, W. of Jennings Road, 7-VII-2005, B. Kondratieff, J. Owens, J. Schmidt, D. Leatherman; Jefferson County, Monticello, 26-IV-2002, on persimmon, R. Mizell; idem, 17-IV-1994, Buck B. Pales; idem, 8-V-1994, Buck B. Pales; idem, 1-V-1994, Buck B. Pales; idem, 2-V-1994, Buck B. Pales; idem, 24- IV-1994, Buck B. Pales; idem, 8-V-1994, Buck B. Pales; idem, 28-V-1994, Buck B. Pales; Duval County, Jacksonville,?-V-?, collection date and collector not noted; Dade County, 1 mi S Florida City, 22-IV-1976, beating miscellaneous slash, F. T. Hovore; Alachua County, 21-V-1978, M. C. Thomas; Alachua County, Payne Prairie State Preserve, 14-16-V-1984, light trap, G. B. Fairchild & H. V. Weems; Alachua County, Austin Cary Forest, 12-21-IV-?- Insect flight trap, Fairchild, & Roberts; idem, 19-21-III-1982, insect flight trap, G. B. Fairchild & H. V. Weems; Alachua County, 9 mi NW Gainesville, University of Florida Horticultural Unit, 16-20-IV-1977, Malaise trap, H. N. Greenbaum; Palm Beach County, Royal Palm Park,?- V-1948; Liberty County, Torreya State Park, 16-V-1971, in blacklight trap, H. W. Weems, Jr., & G. B. Fairchild; Highlands County, Archbald Biological Station, 29-III-1983, hickory, L. L. Lampert, Jr.; idem, 25-IV-1980, L. L. Lampert, Jr.; idem, 4-VI-1978, insect flight trap, H. V. Weems, Jr. & Lisa K. Klein; Putnam County, 11-V-1959, H. W. Weems; Putnam County, Interlachen, 21-III-1953, sweeping oak, B. K. Dozier; idem, 4 mi n Old Town, 18-20-V-1978, E. Giesbert; Dixie County, 4-5 mi N Old Town, 27-IV-1979, beaten from hardwood slash, R. L. Penrose; idem, 5-V-1979, R. L. Penrose; 5 mi N. Old Town, 23-V-1978, on persimmon stump, Woodruff & Choate; Old Town, 30-V-1978, G. B. Edwards; idem, 20-V-1978, G. B. Edwards; Dixie County, 3.5mi N of Old Town, Rt. 349, 13-V-1979, Lloyd R. Davis, Jr.; idem, 27-IV-1979, M. C. Thomas; idem, 8-V-1978, M. C. Thomas; idem, 4 mi N Old Town, 6-7-V-1978, F. Hovore; idem, 11-12-V-1979, E. Giesbert; Seminole County, Econ Wild Area, 17-VI-2000, scrub oak/saw palmetto (burned), Malaise trap, Trevor Smith; 8 miles W Cross City, 30-III-1990, W. F. Chaimberlain; Hernando County, Withlacoochee State Forest, Richloam Tract, 9-IV-1992, M. C. Thomas; idem, 8-IV- 1994, M. C. Thomas; 3.8 mi SW Archer, 17-23.IV.1988, Malaise trap in rosemary turkey oak sand hill, P. Skelley; Volusia County, Port Orange, 5-IV-1962, J. F, Brimley; Monroe County, 3-5-V-1978, F. Hovore; No Name Key Quarry, 12-VI-2013, D. Fine; Liberty County, Torreya State Park, 6-V-1979, M. C. Thomas; idem, 7-V-1989, M. C. Thomas; Bradford County, 8 km W Starke, 23-30-VI-1980, Alan Wilkening; Gadsden County, Aspalaga Landing, Aspalaga Road, 9-V-2005, Gino Nearns; Hillsborough County, University of Southern Florida campus,?-?-1968, collector not noted; idem, 18-VII-1972, collector not noted; Pinellas County, Tarpon Springs, 20-III-1950, H. & H. Howden: Georgia; Jasper County, Berner, 20-VI- 1979, H. Flaschka; Hart County, Neberg, black light, 4-5-VII-1980, F. N. Young; Oglethorpe county, 12-V- 1959, O. Tyre; Baldwin County,?- IV-1973, J, H. West; Berrien County, near Alapaha,?-VI-VII-1973, ex pecan logs, J. R. Ables; Berrien County, 3 mi Alapaha, emerged from wood of pecan, 25-III-1973, R. Turnbow; Dekalb County,?-?-1971, reared, collector not noted; Clinch County,?- V-1973, reared from pecan, collector not noted; Henry County,?-VI-VII-1973, reared from pecan, collector not noted; Walker County, 5 mi SE Villanow, 8-VI-1969, R. L. Wescott; Richmon County, Augusta, 7-V-1946, P. W. Fattig; Fulton County, 28-IV-1943, P. W. Fattig; Barrien County, 3 mi E Alapaha, emerged from wood of pecan during June 1973, R. Turnbow; idem, 2-4-IV-1973, R. Turnbow; Clarke County, Whitehall Forest, 27- III-1973, R. Turnbow; idem, emerged from wood of Cornus florida Linnaeus on 1-6 III-1976, R. Turnbow; idem, 30-VI-1973, R. Turnbow; idem, 30-IV-1972, R. Turnbow; Upson County, Pleasant Grove Road, 1- IV-1993, ex. Persimmon & Ironwood, R. Morris; Lamar County, Barnesville, Howard Road, 3-4-VII- 1999, R. Morris; Tallnall County, 2 mi E 147 Ohoopee River, 15-30-I-1999, limbs of Quercus species , R. Morris; Union County, Blairsville, 18-24-VI-1990, P. J. Landolt; Union County, hwy 180, 4.8 mi NE junction129 & 19, 4-5-VI-1981, R. L. Penrose; Clinch County,?- VI-1973, J. E. Wappes; Twiggs County, Sawmill, 22-V-1970, collector not noted; Green County, Ocanee National Forest, 14-21-V-2004, M. D.Ulyshen: Illinois; Platt County, Allerton Park, 7-VIII-1974, Lloyd R. Davis, Wabash County, Wabash,?- V-1878, F. C. Bowditch; Lake County, Lake Forest, 1-VII-1917, collector not noted; Cook County, Glenview, 2-VII-1905, collector not noted; idem, 4-VI-1915, F. J. Psota; Cook County, Beverly Heights, 15-VI-1913; Cook County, Schiller Park, 21-VII-1935, collector not noted; Cook County, Palos Park, 20- VII-1937, Hank Dybas; Cook County, Glendon Park, 7-VI-1903, A. B. Wolcott; idem, 14-VI-1903, A. B. Wolcott; LaSalle County, Starved Rock State Park, 18-VI-1940, Floyd G. Werner; idem, 23-VI-1940, Floyd G. Werner; Cook County, Edgebrook, 20-VI-20014, E. Liljeblad; Macon County, 12-V-1981, P. Skelley; Mason State Forest, 16-VI-1963, J. M. Campbell; Union County, Pine Hills, 17-V-1963, J. M. Campbell; Monroe County, Bloomington, 6-8-VI-1988, F. N. Young, idem, 4-VII-1898, collector not noted; Coös County, Shelburn,?- VII-1884, collector not noted; La Salle County, Starved Rock State Park, 18-VI- 1940, Floyd G. Werner; Kankakee, County, 19-VI-1937, collector not noted; Winnebago County, Rockford, 13-VI-1944, H. S. Dybas; Du Page County, Glendon Park, 14-VI-1903, A. B. Wolcott; Du Page County, Argonne National Lab 14-VII-1967, J. Wagner & W. Suter; McHenry County, Algonquin, collection day not noted, W. A. Nason; Bond County, Greenville, 26-VI-1966, D. J. & J. N. Knull; Lake County, Ravinia, 21-VII-1935, B. C. Williams; McLean County, Lexington, collection date and collector not noted; Cook County, Riverside, 25-VI-1927, collector not noted; Cook County, Edgebrook, 18-VI-18, F. Psota; Cook County, Glendon Park, 7-VI-1903, A. B. Wolcott; Cook County, Schiller Park, 21-VII-1935, R. Wenzel; Cook County, Chicago, 18-VI-1921, collector not noted; Cook County, Willow Springs, 3-VII-1916, F. J. Psota; idem, 26-VI-1904, W. J. Gerhard; idem, 16-VII-1932, W. J. Gerhard; idem, 30-VI-1912, W. J. Gerhard; idem, 2-VII-1905, W. J. Gerhard; idem, 6-VI-1922, W. J. Gerhard; idem, 4-VI-1905, W. J. Gerhard; idem, 29-VI-1916, collector not noted; idem, 10-VI-1914, collector not noted; idem, 30-VI-1912, W. J. Gerhard; idem, 4-VII-1924, W. J. Gerhard; idem, Willow Springs, 16-VII-1974, C. Selinger; Macon County, 30-VI- 1979, P. Skelley: Indiana; Pulaski County, Jasper-Pulaski Forest, 7-VII-1961, Charles E. White; Clay County, Poland, 6-VI-1963, reared from Carya, Charles E. White ; White County, 12-VII-1961, Charles E. White; Posey County, 19-VI-1963, Charles E. White; Crawford County, 21-VI-1966, Dianne Eckert; Lawrence County, Mitchell, 6-VI-1908; Morgan County, 12-VII-1912, Gerhard; Porter County, Beverly Shores, 16-VII-1935, H. Dybas; Porter County, Dunes State Park, 22-VI-1939, H. Dybas; Lake County, Hessville, 12-VII-1908, W. J. Gerhard; Lake County, 4-VI-1921, Marion County, Indianapolis, 16-V-1938, reared from Elm; idem, emerged from 4 inch branch of elm on 20-VI-1939; Tippecanoe County, Lafayette, 2-I-1998, N. M. Downie; idem, 3-I-1990, M. Downie; idem, 14-VII-1982, M. Downie; idem, 10-VII-1982, M. Downie; idem, 5-VI-1982, M. Downie; idem, 10-VII-1981, M. Downie; Tippecanoe County, McCormick’s Woods, 12-V-1989, N. M. Downie; idem, 24-VII-1989, N. M. Downie; Tippecanoe County, 2-VII-1989, N. M. Downie; idem, 10-VII-1956, N. M. Downie; idem, 4-VI-1962, N. M. Downie; Tippecanoe County, 12- VI-1961, N. M. Downie; idem, 31-V-1951, N. M. Downie; Tippecanoe County, 27-VII-1958, N. M. Downie; Brown County, Bear Wallow, 31-V-1965, Charles E. White; Knox County, 21-VI-1937; Brown County, Bear Wallow, 27-V-1988, J. E. Wappes; idem, 9-VI-1967, Charles E. White; idem, 7-VI-1985, Charles E. White; idem, 12-VI-1967, blacklight trap, Charles E. White; Brown County,?- Vi-1970, N. M. Downie; Brown County, Bear wallow, 10-VI-1978, collector not noted: Idaho; Cassia County, Little Cottonwood Creek, Crater of the moon National Monument, 8-IV-1965, D. S. Horning; Cassia County, Hecker Canyon, winter, 1970, reared from Alder, W. F. Barr; Butts County, 19 mi E Howe, winter 1970, reared from Juniper, W. F. Barr; Lemhl County, Second Creek, Salmon R, George, S. of Salmon, 14-VII-1959, H. B. Leech; Owyhee County, 4 mi W Homedale, 15-VI-1960, R. B. Roberts; Boundary County, Paradise Valley, 23-VI-1960, A. R. Gittins; Canyon County, Parma, 22-VIII-1959, W. F. Barr; idem, 10-V-1954, on peach, 2131 ft, A. J. Walz; idem, 10-V- 1954, 2231 ft, on peach, A. J. Walz; idem, 10-VII-1952, A. J. Walz; idem, Parma, 28-VII-1965, L. de los Reyes: Iowa; Story County, Ames,?-VI-?, collector not noted; Polk County, W Sailorville Lake, 6-9-VI-1985, R. Schieferstein; Howard County, Elma, 10-VII-1962, collector not noted; idem, 10-VII-1904, collector not noted; Dallas County, Waukee, emerged from hickory wood on 13-VI- 1972, A. Stewart Cott; Linn County, 3.0 mi N Viola, 17-22-VII-1972, E. L. Sleeper: Kansas; Woodson County, 3 miles SW Piqua,?- IV-1996, D.J. Heffern; Atchison County, Atchison, 10-VI-1956, J. W. McReynolds; Labette County, Big Hill Reservoir, NW of dam, 37°16'.55'’ N 95°28'.43'’W, 13-28-VI-2005, G. A. Salsbury, Cherokee County, 2 mi S Galena, 37°02'.64'’ N 94°38'.18'’W, 13-V-2-VI-2005, Canopy trap, G. A. Salsbury; idem, 13- 28-V-2-VI-2005, Canopy trap, G. A. Salsbury; 14-V-2-VI-2005, Canopy trap, G. A. Salsbury; Crawford County, 3 mi NE Pittsburg, NW of dam, 37°26'.64'’ N 94°37'.04'’W, 13-VI-2005, Canopy trap, G. A. Salsbury; Jefferson County, The Falin property, 1.5 km N jct.94 th Sreet & Kingman Road, 39°13'.38'’ N 95°24'.24'’W, 20-27-VI-2005, ex Malaise trap, lower meadows, Z. H. Falin; idem, 24- 29-VI-2005, ex Malaise trap, lower meadows, Z. H. Falin; 10-20-VI-2005, ex Malaise trap, lower meadows, Z. H. Falin; 29-V-7-VI-2005, ex Canopy Malaise trap, lower meadows, Z. H. Falin; 20- 19-24-V-2005, ex Canopy Malaise trap, lower meadows, Z. H. Falin; 27-VI-3-VII-2005, ex Canopy Malaise trap, lower meadows, Z. H. Falin; Jefferson County, Perry State Park, 4-IV-1977, ultraviolet light, G.H. Nelson; Shawnee County, Topeka, 16-VI-1904, Knaus; Barber County, Medicine Lodge, 2-VI-1975, on Sapindus drummondii Hook & Arn. ; Reno County, 6-VI-1914, J. Warren; Pottawatomie County, Onaga, 10-VI- 1901, Crevencoon; Pottawatomie County, St. Mary, 4-VII-1938, J. W. Green; Saline County, Salina,?- V- 2000, W. Opitz; Riley County, Kings Creek, Konza Prarie Nature Area, 20-V-1986, Baumann & Nelson; Riley County, 24-VI-?, Popenoe; Riley County, Upper King Creek, Konsa Prarie Natural Area, 20-V- 1986, R. W. Bauman; 12-VI-1978, S. Krueger; Riley County,?- VII-1955, V. E. Adler; Jefferson County, Perry State Park, 3-VI-1977, UV light, G. H. Nelson; Douglas County, 2 mi NW of Baldwin, between 1550E & 1600E Roads at 500N Road, 38°48'.58'’ N 95°12'.56'’W, 9-16-VI-2005, Canopy Trap, S. White; Douglas County, Lawrence, 18-VI-1922, C. H. Curran; Chase County, 21-VI- 1923, W. J. Brown; Johnson County, Shawnee, 1.2 mi N of 43 rd Street, near Kansas River, 39°02'.82'’ N 94°48'.94'’W, 2-9-VI-2005, ex Canopy trap, S. White: Kentucky; Green County, Crail Hope, i-VII-1946, J. W. Green; idem, 13-V-1946, J. W. Green; Christian County, 26-VI- 1961 m J. M. Campbell: Louisiana; East Feliciana Parish, Thompson Creek, 1 miles W Jackson, 1-V-2002, H. & A. Howden; Saint John The Baptist Parish, Edgard, 18-V- 1973, Ultra violet light, V. Brou; Iberville Parish, Sunshine, 5-VIII-1972, V. A. Broun; Winn Parish, 21- VI-1915, collector not noted; Winnfield,?- VI-1916, collector not noted; Acadia Parish, Crowley, 13-IV- 1980, F. H. Shoemaker; Rapides Parish, Alexandria, 6-V-1963, light trap, L. A. Cambre; Natchitoches Parish, Campti, 22-VII-1978, S. M. Clark; Claiborne Parish, Corney Lake 27-I-1983, B. P. & J. L. Carr; Baton Rouge Parish, Place DuPlantier Apts., 27-IV-1985, E. G. Riley; La Bossier Parish, 11 km NE Bossier City 15-VI-2013, black light, Brandon Hays: Maine; Oxford County, Paris, 10-VII-1915, C. A. Frost; Kennebec County, Monmouth, 27-VI-1912; idem, 26-VI-1905, collector not noted: Maryland; Dorchester County, Cambridge, 14-VI-1947, H. L. Dozier; Baltimore County, 5-VII-1931, J. W. Green; Prince George’s County, Laurel, 25-VI-1965, Malaise trap; Prince George’s County, United States Agricultural Center, 27-VI-1954, fresh break in gum sapling, H. L. Dozier; Prince George’s County, Hyattsville, 5-VI-1919, L. L. Buchanan; idem, 19-V-1918, L. L. Buchanan; Prince George’s County, College Park, 23- VI-1951, oak slash, B. K. Dozier; idem, 13-VI-1934, oak slash, B. K. Dozier; idem, 28-V-1967, on foliage of black locust, B. K. Dozier; Prince George’s County, Beltsville, 27-VI-1934, oak slash, B. K. Dozier; Frederick County, Braddock Heights, 14-VI-1952, on slash, B. K. Dozier; Charles County, nanjemoy 3- VI-1948, H. Howden: Massachusetts; Barnstable County, Barnstable,?- VII-1898. Collector not noted; Bristol County, 2-VIII-1923, N. S. Easton; Plymouth County, 29-V-1932, C. E. White; Suffolk County, Dorchester, 9-VI-1907, collector not noted; Suffolk County, 8-VII-1915, Boister; Bristol County, 15-V- 1915, N. S. Easton; Hampshire County, Notch Roas, S, Amherst, 24-VI-1998, F. Knab; Norfolk County, Brookline, collection date not noted, Bowditch; idem, 22-VI-1901; Norfolk County, 25-VI-1910, Bolster; Norfolk County, Brooklyn, D. A. Fenyes; Norfolk County, collection date not noted, Bowdich; Norfolk County, Dover, 20-VI-1916, F. W. Graham; Norfolk County, 11-XI-1899, collector not noted; Middlesex County, Framingham, 4-VIII-1920, C. A. Frost; Middlesex County, Newton,?-VI-?, collector not noted; Middlesex County, Cambridge, collection day and collector not noted; Middlesex County, 7-VI-1951, W. Rosenberg; Northampton, 13-VII-1971, E. J. Kiteley; Tyngsborough, 27-VII-1873, collector not noted: Michigan; Clinton County, Rose Lake, Wild. Reserve area, 4-VII-1987, sweeping vegetation, J. Jenkins; Saint Joseph County, vicinity Klinger Lake, 23-VI-1975, E. Giesbert; Livingston County, 2 mi N Whitmore Lake, 14-VII-1956, beating Quercus alba, G. H. Nelson ; idem, 26-VI-1956, on trunk of dying Populus tremuloides eating Agrilus, G. H. Nelson ; Washtenaw County, Ann Arbor, 4-XII-1968, in wood cellar, W. W. Newcomb; Marquette County,?- VI-1928, Collector not noted; Presque Isle County, Ocqueoe State Park, 5-VII-1961, K. Willis; Berrien County, Harbert, 27-VII-1974, E. Giesbert; idem, 23-VI-1985, E. Giesbert; Cheboygan County, 16-VII-1945, Alyce Ridinger: Minnesota; Itasca State Park, 21-VII-1937, H. R. Dodge; Crow Wing County, Brainerd, 12-VI-1962, E. J. Kiteley; Clearwater County, Itasca State Park, June-?-1957; Butler County, 16-VI-1961; Anoka County, Brine Road, 25-VI-1951, E. J.Kiteley; Hubbard County, Park Rapids, 4-VII-1959, E. J. Kiteley; Cook County, Gunflint Trail Road 2-VII-1984, slash areas/log depots, Downie & Wappes; Saint Louis County, 14 miles W Ely, 30-VI-1984, Wappes & Downie: Mississippi; Harrison County, Gulfport, 21-VI-1968, collector not noted; Washington County, Stoneville, 10-11-VI-1981, F. Hovore; Washington County, Carroll County, 2 mi W Carrollton, 29-V-1981, F. Hovore; Scott County, Forkvillem 1 mi S, 9-VI-1981, R. L. Penrose; Montgomery County, 3.0 mi W Stewart on hwy 82, 22-VI-1999, mixed hardwood slash; Lafayette County, 19-VI-1954, F. M. Hull: Missouri; Green County, 11-18-V-2014, Rogersville, 8725 E Buckhorn Lane, J. Courtney, K. Smith, S. Fitzgerald; Henry County, 7 mi E Clinton, 17-VI-1973, on Diospyros virginiana Linnaeus ; Vernon County, 4-VII-1969, McReynolds; Saint Louis County, Clarkson Valley, Jct Orville & Shepard Rds, emerged 24- 31-V-1998 ex dead branch of Juglans negra L., T. C. MacRae; Saint Louis County, Chesterfield Village, Monsanto Campus, emerged 1-16-V-1999 ex dead branch of Acer saccharum Marsh, T. C. MacRae ; Ripley County, Bald Hill Glade National Ar, 4.6 mi SW hwy V, emerged 17-31-VII-1997 ex trunk of Quercus stellata, T. C. MacRae ; Berry County, 3.2 mi S Roaring River on hwy 112, 14-VI-1997, on Carya species , T. C. MacRae; Taney County, 2 mi N Forsyth, 8-V-2000, Eric Eaton; Taney County, Mincy Conservation Area, 27-VI-1998, UV light in oak/hickory forest, T. C. MacRae; idem, emerged from dead branch of Quercus muhlenbergi during 9-15-VI-2001, T. C. MacRae; idem, emerged from dead branch of Quercus muhlenbergi during 16-22-VI-2001, T. C. MacRae; Dade County, Stockton Lake, emerged 1-7-VI-1998, ex trunk of Carya ovata (P Mill.) K Kotch, T. C. MacRae ; Carter County, Pinewoods Lake National Forest, 2-VI-1984, deciduous forest, shallow lake and pinewoods, blacklight, J. R. Heiteman; Mark Twain National Forest, Pinewoods Lake, 2.0 mi W Ellsmore, emerged from cut log of Carya during 17-31-V- 1999, T. C. MacRae; Stoddard County, Holly Ridge State Forest, emerged 8-14-VI-1989, dead 1-3'’ branch Prunus serotina Ehrh. T. C. MacRae ; Bollinger County, Duck Creek Cons. Area, NE Corner Pool 2, emerged from upper trunk/main branch of dead Carya alba (L.) Nutt. During 24-31-V-1998, T. C. MacRae; Oregon County, Mark Twain National Forest, McCormack Lake, 3-VI-1976, on Juglans negra L., G. H. Nelson; idem, Junction hwy 19 & Eleven Point R, 3-VI-1976, on dead trunck of Quercus falcata, G. H. Nelson ; Butler County, 2.4 mi E hwy W University Forest, 4-VI-1999, on cut stump of Quercus coccinea Muenchh, T. C. MacRae ; Jasper County, Carterville, 27-VII-1986, Gregory V. Myers; Kansas City; Jackson County, Lees Summit, 18-VII-1947, R. E. White; Burr Oak Woods, Natural Area, 24-VI-1998, on Quercus macrocarpa “en copula”, G. H. Nelson;,?- VII-1953, Ronald H. Pine; Blue Springs, 7-VII-2000, downed limb of Robinia pseudo-acacia, G. H. Nelson ; idem, 20-V-1998, G. H. Nelson; idem, 28-VI-2000, G. H. Nelson; idem, emerged from wood of Gleditsia triacanthos (Linnaeus) , G. H. Nelson; idem, 5-VI- 1997, G. H. Nelson; idem, 7-VII-2000, on Gleditsia triacanthos (Linnaeus) ; idem, 29-VI-1953, D. J. & J. N. Knull; idem, 13-IV-1955, E. J. Kitelley; Valeey View glades, 13-V-1987, T. C. MacRae; Raytown, 14- VII-1970; Warren County, 10 mi W Highway F, Village of Insbrook, 3-VII-1999, T. C. MacRae; idem, emerged from the wood of Ulmus americana on 5-VI-1970, G. H. Nelson; idem, 7-VI-1969, on Cercis canadensis, G. H. Nelson ; idem, emerged from wood of Ulmus americana on 22-XII-1968, G. H. Nelson; idem, 7-VIII-1970, G. H. Nelson; idem, 10-VI-1970, G. H. Nelson; idem, 30-VI-1978, ultraviolet light, G. H. Nelson; idem, 23-VI-1974, on Carya ovata, G. H. Nelson ; idem, 25-VI-1974, on Carya ovata, G. H. Nelson ; Wayne County, Williamsville, 7-VII-1955, U V light trap, E. C. Becker; Carter County, Peck Ranch Conservation Area, Stegall Mountain, 16-23-V-2001, on lower trunk of fire-killed Quercus velutina, T. C. MacRae ; idem, emerged from dead branch of Amelanchier arborea during 24-31-V-2001 ,, T. C. MacRae; Saline County, Nelson, 2-VI-1974, E. G. Riley; Miami Access, emerged from dead branch of Quercus macrocarpa Michx. , during 25-30-VI-2002, T. C. MacRae; Adaire County, Kirksville, 13-VI-1981, M. Huybenez; Pike County, New Hartford, 13-VI-1940, W. B. Craig, Clay County, 2 mi W Missouri City, Coolie Lake, 2-VII-1997, on Acer negundo, G. H. Nelson ; idem, 22-VI-1997, on Acer negundo, G. H. Nelson ; Boone County, 25-VII-1967, S. Poe; Boone County, Ashland Wildlife Ar. 17-VI-1977, E. G. Riley; Randolph County, 1 mile E Moberly, 9-VI-1974, E. G. Riley; Randolph County, Sugar Creek lake, 15-VI- 1974, E. G. Riley; Stone County, 3 miles NE Blue Eye, 12-V-1987, T. C. MacRae; Franklin County, Meramec State park, 15-VI-1987, T. C. MaCrae; Boone County, 15-VI-1973, light trap, J. W, Smith: Montana; Missoula County,?-?-1913, C. C. Adams: New Mexico; Chaves County, 27-VII-2009, J. Hardin: Nebraska; Douglas County, 30-VI-1938, D. J. & J. N. Knull; Valley, 30-VI-1938, D. J. & J. N. Knull: New Hampshire; Carroll County, Chocorua, collection day and collector not noted; Rockinham County, 10- VII-1901, S. A. Shaw Crafton County, Hanoverm?- VI-1928, collector not noted: New Jersey; Gloucester County, Paulabero, 11-VI-1961, H. L. Dozier; Camden County, Camden,?- VI-1917, collector not noted; Greenwood Lake, 20-VI-1924, collector not noted; Burlington County, Medford, 2-VII-1939, E. J. F.Marx; Burlington County, Atsion, 12-VI-1945, J. W. Green; idem, 28-VI-1946, J. W. Green; Ocean County, Lakehurst, 25-V-1960, W. F. Barr; Orange County, Fort Montgomery, 3-VII-1915; Somerset County, Griggstown, 8-VI-1936, reared from Elm, C. H. Hoffmann; Orange County, Greenwood Lake, 7-VI-1942, M. Cazier; Essex County, South Orange, 16-VI-1889; Essex County, Newark, collection date and collector not noted; Essex County, Montclair, 25-VI-1922, A. Nicolay; Essex County, Montclair, 19-VI-1925, A. Nicolay; Essex County, South Orange, 8-VI-1959, collector not noted; idem, 30-VI-1889, collector not noted; Essex County, Bloomfield, 20-VI-1939, feeding on Scolytus multistriatus Marsh. ; Mercer County, Princeton, 4-VI-1934, bred from Elm, Woodley; idem, 9-VI-6-VII-1943, collector not noted; Camden County, Camden; Camden County, Clementon,?- V-1930, collector not noted; Cape May County, 21-VI- 1941, J. W. Green; Cape May County, Ocean City, 21-VI-1941, R. C. Casselberry; Warren County, 1-VII- 1917, J. W. Green; Cumberland County, Rutgers experiment Station, 21-V-1988, J. E. Wappes; Bergen County, Passaic Junction 7-VII-1912, collector not noted; Bergen County, Fort Lee, collection date and collector not noted; Monmouth County, 16-VI-1935; Somerset County, Somerville,?-VI-?, Schott: New York; Columbia County, Mellenville, 10-VII-2007, Lindgrin funnel, J. Dallessandro; Saint Lawrence County, Canton, 4-VI-1935, N. M. Downie; Orange County, West Point, 9-VI-1915, W. Robinson; Orange County, Fort Montgomery, 3-VII-1915, F. M. Shoft; Rockland County, 12-VI-?, collector not noted; Queens County, Queens, 31-V-1924, F. M. Schott; Tompkins County, Six Mile Creek, 9-VII-1959, collector not noted; Tompkins County, Ithaca,?- VII-1917, collector not noted; Westchester County, Peekskill, collection date and collector not noted; idem, collection date and collector not noted; idem, 15-VI-1889, Sherman; Suffolk County, Wading River, 23-VI-1917, collector not noted; Onondaga County, 14-VII-1946, N. M. Downie; Suffolk County, Montauk, Long Island, 4-VII-1916, F. M. Schott; Onondaga County, Syracuse, collection date and collector not noted; Onondaga County, 21-VI-1941, N. M. Downie; Nassau County, Roslyn, Long Island, 24-VI-1930, J. N. Belkin; Tomkins County, 10-VIII-1940, J. N. Belkin; Kings County, Brooklyn, Flatbush, 3-VI-1896, J. L. Zabriskie: North Carolina; Rockingham County, Mayo River State Park,?-IV-V-2015, flight intercept trap, W. D. Merritt; Rockinham County, Reidsville, 30-V-1973, J. S. Ashe; Buncombe, Black Mountain, 1-VI-1912, Beutenmuller; Anson County, Peachland, 15-VI-1953, oak slash, B. K. Dozier; Wake County, Raleigh, 2-VI-1952, B. K. Dozier; Moore County, Southern Pines, 24-V-?, A. H. Manee; Jackson County, Highlands, Whitesides Mountain, 1-VII-1957, W. J. Brown; Jackson County, 19-VI-1959, W. Rosenberg; Orange County, University of North Carolina, 30-V-1972, Wharton; Burke County, Lake James, State Park, 13-V-26-VI-2014, flight intercept trap, W. D. Merritt; Cumberland County, Cavers Creek State Park,?-VII-VIII-2011, flight intercept trap, W. D. Merritt; Guilford County, Haw River State Park, V-VI-2015, flight intercept trap, W. D. Merritt; Camden County, Dismal Swamp State Park, 2-V-7-VI-2011, flight intercept trap, W. D. Merritt; Mecklenburg County, Charlotte, VI-VII- 1998, ultra violet light trap, J. F. Cornell; Wake County, 7 miles SW Raleigh, off road 1152, 1-VI-1985, Malaise trap, C. S. Parron; Chovan County, 1.6 km W Edentun, road 1200, 18-VI-1987, A. C. Evenson, Jr: North Dakota; Ransom County, 5 mi NW McLeod, 4-VII-1968, H. F. Howden: Ohio; Clermont County, 28-V-1962, R. D. Mathis; Champaign County, 6-VI-1954, R.E. Woodruff; Green County Ohio, 14- V-?, D. J. & J. N. Knull; idem, 13-VI-1956, D. J. & J. N. Knull; idem, 22-VI-?, D. J. & J. N. Knull; Delaware County, 27-VI-1943, D. J. & J. N. Knull; idem, 8-VII-1942, D. J. & J. N. Knull; idem, 17-VII-1945, D. J. & J. N. Knull; idem, 22-VI-?, D. J. & J. N. Knull; Green County, 15-VI-1953, D. J. & J. N. Knull; Scioto County, 9-VI-1945, D. J. & J. N. Knull; Hocking County, 29-V-?, D. J. & J. N. Knull; idem, 16-VI-1959, J. N. Knull; idem, 14-VI-1950, D. J. & J. N. Knull; 1-VI-?, D. J. & J. N. Knull; Hamilton County, Cincinnatti, Lower Price Hill, 23-VI-1996, E. Eaton; Hamilton County, Cincinnati, 23-VI-1903, collector not noted; idem, Clifton, 13-VI-1990, Eric Eaton; Franklin County,?- VII-1927, J. N. Knull; idem, 9-VI-1952, D. J. & J. N. Knull; Coshocton County, Cavallo; Guernsey County, Millwood, 19-VII-1940; Guernsey County, 19- VII-1940; Knox County, 5-VI- 1942, H. F. Strohecker; Chillicothe, 10-VI-1923, A. E. Miller; Stoney Creek Forest, 6-14-1928; Vinton County, Lake Hope state park, 25-V-1985, R. Androw; Muskingum County, Zanesville, 24-VI-1987, P. W. Kovarik: Oklahoma; Comanche County, Fort Sill, East Range, 11-13-VI- 2002, B. Condratieff, J. Scmidt,, D. Leatherman; Garfield County, Fairmont, 9-12-VII-1982, John F. Reinert; idem, 9-11-VII-1980, insect flight trap, John F. Reinert; Cherokee County, Tahlequah, 170 VI- 1939, Kaiser-Nailon; Marshall County, Willis, 1 mi SE, 6-VI-1972, W. Suter; Marshall County, Lake Texoma, 13-VI-1969, W. Suter; Pontotoc County, Ada, 17-Vi-1952, D. J. 7 J. N. Knull; Wichita National Forest, 9-VI-1932, E. B. Webster; Cleveland County, Norman,?- VI-1980, Willian D, Shepard; Cleveland County, 11-VI-1976, light trap, William D. Shepard; idem, 30-VII-1976, William D. Shepard; Muskogee County, 3mi W Webbers Falls, 7-VI-1964, H. S. Dybas; Muskogee County, Camp Gruber, 6-VI-1943, J. A. Wilcox; Marshal County, Oklahoma University Biological Station, 24-VII- 1969, R. H. Arnett, Jr.; Oklahoma City, summer 1958, W. Rosenberg; Hayes County, Chouteau, 27-VI-1967, on weed, D, C. Arnold; Osage County, Ponca Lake, 10-VII-1981, K. Burnham; Latimer County,?- V-1986, K. Stephan; idem,?- IV-1987, K. Stephan; idem,?- V-1987, K. Stephan; idem,?- VII-1986, K. Stephan; idem,?- IV-1986, K. Stephan; idem,?- VI-1987, K. Stephan; idem,?- IV-1988, K. Stephan; idem,?- VI-1988, K. Stephan; idem,?- V-1988, K. Stephan; idem,?- V-1989, K. Stephan; idem,?- VI-1989, K. Stephan; idem,?- VI-1991, K. Stephan; idem,?- VII-1991, K. Stephan; idem,?- X-1988, K. Stephan; idem,?- IV-1986, K. Stephan; idem,?- VII-1988, K. Stephan; idem,?- V-1982, K. Stephan; idem, VI-1989, Karl Stephan; Latimer County, 5 mi W Red Oak,?- VII-1981, K. Stephan; idem, 11-VII-1977, K. Stephan; idem, 11-VII-1977, K. Stephan; idem, 7-V-1977, K. Stephan; idem, 11-VI-1977, K. Stephan; idem, 30-IV-1977, K. Stephan; idem,?- VI- 1984, K. Stephan; idem,?- V-1984, K. Stephan; idem,?- VI-1985, K. Stephan; idem,?- VII-1983, K. Stephan; idem,?- VI-1983, K. Stephan; idem,?- IV-1985, K. Stephan; idem,?- VII-1982, K. Stephan; idem, 2-VII- 1977, Karl H. Stephan; Payne County, VI- 1924, W. J. Browa; Payne County, Stillwater, 28-V-1981, D. C. Arnold; LeFlore County, Lake Wister State Park,?- V-1992, ex. Quercus, D. J. Heffern : Oregon; Deschutes County, Cline Falls, 8-VII-1946, Frank M. Beer; Baker County, Wallowa Mountains, 4-VII-1922, collector not noted; Umatilla County, 11.5 mi W Rieth, emerged from wood of Betula occidentalis on 27-X- 1972, R. L. Westcott; Pennsylvania; Fulton County, 1-2 mi N Covans Gap State Park, 10-11-VII-1978, E. Giesbert; Huntingdon County, Huntingdon, State Game Lands # 112B, M Gap Road, 5-VI-2001, Frank D, Fee; Huntingdon County, 2 mi E Neelyton, 18-VI-1989, J. E. Wappes; Lancaster County, collection date and collector not noted; berks County, Mount Penn, 20-VI-1950, D, C. Kissinger; Berks County, Stoney Creek Mills, 26-VII-1950, D. G. Kissinger; Berks County, Stoney Creek Mills, 12-VII-1950, D. G. Kissinger; Berks County, Mount Penn, 26-VI-1950, D. G. Kissinger; Franklin County, Mount Alto, 5-VII-?, J. N. Knull; Allegheny County, collection date and collector not noted; Dauphin County, Linglestown, 24-VI-?, collector not noted; Dauphin County, Clarks Valley, reared, collection date not noted, J. N. Knull; idem, 15-VI-1958, J. N. Knull; Tioga County, 21-VI-1918, collector not noted; Cumberland County, Lemoyne, 23-IV-1910, collector not noted; idem, 3-V-1910, J. N. Knull; Cumberland County, 15-V-1911, collector not noted; Cumberland County, Lemoyne, 19-V-1910; idem, 5-X-1910, H. H. Kirk; Dauphin County, Harrisburg, 14-IV-1911, H. B. Kirk; idem, 22-V-1922, A. B. Champlain; idem, 14-IV-1911, H. B. Kirk; Dauphin County, Harrisburg, 21-IV-1911, emerged from elm, H. H. Kirk; Dauphin County, Hummelstown, idem, 26-VI-1933, J. N. Knull; idem, 4-VII-1915, J. N. Knull; idem, 26-VI-1933, J. N. Knull; idem, 3-V-1912, J. N. Knull; idem, 26-VI-1923, J. N. Knull; idem, 30-VI-1912, J. N. Knull; idem, 1- VI-1933, J. N. Knull; idem, 2-VI-1920, J. N. Knull; Franklin County, Mount Alto, 4-VII-?, J. N. Knull; Delaware County, Glenolden,?- V-1912, collector not noted; idem, 12-V-?, collector not noted; Delaware County, 6-V-1904, J. W. Green; Northampton County, 14-VII-1931, J. W. Green; Delaware County,?- VI- 1918, collector not noted; Northampton County, 18-VI-1923, J. W. Green; idem, 5-VIII-1912, W. Green; Philadelphia County, Philadelphia, collection date and collector not noted; Philadelphia County, Angora, 18-VI-?, W. Green; idem, 20-VI-?, W. Green; idem,?- VI-1915;Chester County, Downingtown, 4-VII- 1935, L. J. Bottimer; Abington County,?- VI-1920, collector not noted; Montgomery County, Abington, collection date and collector not noted; NE Jamison, Horseshoe bend, Neshaminy Greek,?- VIII-1956, W. Ivie; Monroe County, Delaware Water Gap, collection date and collector not noted: Rhode Island; Providence County, Providence, 18-VII-1926: South Carolina; Pickens County, Table Rock,?- VIII- 1958, D. Thornton; Clemson, 13-V-1989, J. K. Moulton; Dovehaven, 7 mi NE of Pickens, 19-VII-1990, H. L. Dozier; idem, 20-V-1984, H. L. Dozier; idem, 7-V-1979, H. L. Dozier; idem, 23-V-1989, H. L. Dozier; idem, 6-VI-1983, Annie Dozier; Cherokee County, Kings Mountain National Park, 23-V-1973, L. L. Lambert; Oconee County, CCC camp F2, 30-VI-1036, O. L. Cartwright: Tennessee; Hamilton County, Signal Mountain, 10-V-1922, C. F. Dalman. Warren County, McMinnville, 20-VI-2003, sticky trap, J. B. Oliver; idem, 24-VI-2003, J. B. Oliver; idem, reared from Quercus during 2004-2005, Nadeer N. Yousef; Lake County, Reelfoot, Lake, 2-VI-1954; Franklin County, 7-VI-1958, at light, J. A. Steeves; Morgan County, Deer Lodge,?- VI-1937, Bernard Benesh; idem, 22-VI-1932, Bernard Benesh; Morgan County, Burrville, B. Benesh; Cannon County, Woodbury, 23-V-1993, E. J. Ford; Knox County, Knoxville, 10-V- 1955, H. &. H. Howden; Blount County, Chilhowee Mountain 21-VI-1954, on oak, H. & A. Howden: Texas; Kelley’s Pond, Sam Houston Notional Forest, 15-VI-1990, W. F. Chamberlain; Sabine County, 8 miles E Hemphill, 7-V-4-VI-2011, D. Heffern & B. Raber; Lee County, 14-VI-1907, collector not noted; Fedor, 6-VI-1904, Birkmann; Matagorda County, 2 mi SE Blessing, 19-IV-1984, Marlin Rice; Wharton County, 8-V-1963, Macay, black light trap, Marlin Rice; 8 miles NW El campo, 19-IV-1986, D. J. Heffern; Montgomery County, The Woodlands, 12-13-V-1978, N. M. Downie; Robertson County, 8 miles E Hearns, 21-27-IV-1991, M. Hallmark; Benchley, 30-IV-1941, D. J. & J. N. Knull; Cameron County, Brownsville, Esperanza Ranch, 1-VIII-?, collector not noted; Kleberg County, Kingsville, collection date not noted, C. T. Reed; Lee County, Fedor, collection date not noted, Bock; Nacogdoches County, near Garrison Camp, Whispering Pines, 28-VI-1981, N. Rulien; Nacogdoches, 7-IV-1968, H. R. Burke; idem, 25-VI-1965, at electric light, R. R. Murray; idem, 20-IV-1971, at electric light, William D. Shepard; idem, 4-VIII-1966, fresh cut oak log, R. R. Murray; Parker County, Weatherford, 30-I-1974, Erik L. Klee; Anderson County, Gus Engeling Wildlife Management Area, 2-V-1999, S. M. Clark; Dallas County, 25-V-1907; Dallas County, 12-VI-1938, J. H. Robinson; Dallas, 15-V-1907, F. C. Prott; Brazos County, Near Millican, 12-IV-1986, J. E. Wappes; College Station, 20-VIII-1950, L. S. & E. S. Dillon; idem, 31-V-1919, H. J. Reinhard; Brazos County, nr Millican, 12-IV-1986, J. E. Wappes; College station, Lick Dreek Park 15-30-VI-2000, yellow cup trap on dead Quercus stellata, E. G. Riley ; 1.5 W Bryan, 19-25-VI-1982, R. Turnbow; 10 mi S College Station, 22-26-IV-1991, R. Turnbow; idem, 5-25-1981, beaten from dead oak, R. L. Penrose; idem, hwy 30, Navasota River, 21-IV-1988, R. Anderson; Grayson County, 18-VI-1983, B. F. & J. L. Carr; Jones County, 12 mi W Anson, 2-VI-1982, Cicero; Harris County, Houston Airport, emerged 20-31-1991, D. J. Heffern; Harris County, near W. Houston Airport,?- IV-1994, reared from dead Zanthozylum clavaherculis, D. J. Heffern; Fort Bend County, Brazos State Bend Park, 21-IV-1998, D. J. Heffern; Montague County, Forestburg, 10-VII-1948, collector not noted; Montague County, S Shore Lake Nocona, 26-IV- 2003, J. Owens, J. Scmidt; Terran County, Doug Russel Park, 17-VII-1982, C. S. Wolfe; Angelina County, Angelina National Forest, 5.5 miles SE Zavalla, 19-IV-2-V-1996, Clark, Menard, & Riley; Erath County, Stephenville, 6-XI-1980, T. Riherd; Dickens County, White River Reservation, 10-19-VI-1988, R. Morris; Crocket County, 28 miles E Iraan, 1-VI-1973, Gaumer & Clark; Dimmit County, 13-III-1976, S. E. Jones, Smith County, 27-III-1956, collector not noted; Kerr County, Kerrville, 20-VI-1996, W. F. Chamberlain; Jackson County, 4 miles SW Francitas,?- VI-1992, emerged from Ulmus crassifolia, D. J. Heffern ; Baxar County, 19-IV-1932, H. B. Parks; Liberty County, Liberty, 15-IV-1934, collector not noted; Madison County, 17-VII-1932, collector not noted; Cherokee County, 5-V-1952, H. J. Reihard; Wood County, N Hawkins, 20-V-1998, C. Wolfe & O. Marqua; Kaufman County, 10.4 km NE Forney, 22-VI-2013, Brandon Hays; Milam County, 5.6 km NE Gause, 27-V-1995, H. & A. Howden: Virginia; Rockbridge County, Vesuvius, 25-VI-1956, B. & B. Valentine; Campbell County, Lynchburg, 12-V-1990, J. R. Mawdsley; idem, 13-VI- 1987; Northwest Virginia, 8-V-1952, G. H. Nelson; Fairfax County, East Falls Church, 4-VI-1916, J. N. Knull; Fairfax County, 22-VI-1923, A. Nicolay; idem, 4-VI-1916, J. N. Knull; Blue Ridge Parkway, milepost 100, 21-VI-1988, N. M. Downie; Hampton, 12-V-1944, N. m. Downie; Boutetourt County, Highway 615, near Hipes, 10.5 road mile W Junction Highway 220, 24-V-2006, E. G. Riley; Appomattox County, Vera, 21-VI-1984, Pierre Bélanger; Fairfax County, Alexandria, collection date and collector not noted:

West Virginia; Richland County, Lone Rock , 27-VII-1906 ; Greenbrier County, White Sulphur Springs , 3-VII-1912, W. Robinson. Wisconsin ; Dane County, 10-VII-1940, H. R. Dodge ; Walworth County, Delavan ,?- VIII-1962, Eugene Ray ; Walworth County, 30-VI-1940, H. S. Dybas ; Polk County, Balsam Lake , 1-VIII- 1958, E. D. Kiteley. “Tymgo” 27-VII-1873 ; Monitowoc County, Point Beach State Park , 20-VII-1966, H. Darling. Specimens are deposited in: AMNH, BYUC, CASC, CMNC, CNCI, CSUC, FSCA, MCZC, NCSU, PMNH, SEMC, TAMU, UGCA, USNM, WFBM, and WOPC .

Figures

Figure 66. Geographical distribution of Chariessa pilosa.

Figures 1-18. Antennae of Chariessa spp. 1) Chariessa duponti (male). 2) C. duponti (female). 3) C. floridana (male). 4) C. floridana (female). 5) C. catalina (male). 6) C. catalina (female). 7) C. elegans (male). 8) C. elegans (female). 9) C. texana (male). 10) C. dichroa (male). 11) C. dichroa (female). 12) C. pilosa (male). 13) C. pilosa (female). 14) C. ramicornis (male). 15) C. ramicornis (female). 16) C. vestita (male). 17) C. vestita (female). 18) C. texana (female).

Figures 21-29. Pronota of Chariessa spp. 21) Chariessa catalina. 22) C. elegans. 23) C. ramicornis. 24) C. duponti. 25) C. floridana. 26) C. texana. 27) C. dichroa. 28) C. pilosa. 29) C. vestita.

Figures 30-35. Pronota and elytra. 30-31. Pronota. 30) Chariessa floridana. 31) C. pilosa. 32-35. Elytra. 32) C. ramicornis. 33) C. texana. 34) C. pilosa. 35) Pelonium lampyroides.

Figures 45-48. Habitus of Chariessa spp. 45) Chariessa floridana. 46) C. pilosa. 47) C. pilosa. 48) C. ramicornis.

Figures 57-62. Male genitalia and aedeagi. 57) Chariessa catalina. 58) C. dichroa. 59) C. duponti. 60) C. elegans. 61) C. floridana. 62) C. pilosa.