Ochthephilus rosenhaueri ( Kiesenwetter, 1850 )

Makranczy, György, 2014, Revision of the genus Ochthephilus Mulsant & Rey, 1856 (Coleoptera: Staphylinidae, Oxytelinae), Revue suisse de Zoologie 121 (4), pp. 457-694 : 548-555

Ochthephilus rosenhaueri ( Kiesenwetter, 1850) Figs 28, 212-216, 231, 527, 572

Trogophloeus rosenhaueri Kiesenwetter, 1850: 220 . – Kiesenwetter, 1851: 428.

Ancyrophorus rosenhaueri (Kiesenwetter) . – Kraatz, 1857: 887. – Ganglbauer, 1895: 668. Ochthephilus rosenhaueri (Kiesenwetter) . – Herman, 1970: 385. – Makranczy, 2001: 181. Trogophloeus longipennis Fairmaire & Laboulbène, 1856: 614 .*

Ancyrophorus jailensis Scheerpeltz, 1950: 57 , 66, syn. nov.

Ancyrophorus gracilis Fagel, 1951a: 6 . – Fagel, 1968: 192

Ancyrophorus corsicus Fagel, 1956: 58 , syn. nov.

*starting with Kraatz, 1857 the use of the name ' longipennis ' is totally mixed up, and such confusing references are omitted here

TYPE MATERIAL EXAMINED: Trogophloeus rosenhaueri – LECTOTYPE (here designated): “Kiesenwetter \ Sammlung; Cl. Müller \ Rosenhau-; eri; Kiesw. \ Sammlung; Clemens Müller \ Tirolis [Tirol *+47.00/+11.00*]; D. Rosenhauer \ Lectotypus; Trogophloeus; rosenhaueri Kiesenwetter ; [on the back] des. Makranczy, 1999 \ Ochthephilus ; rosenhaueri Kiesenwetter ; det. Makranczy, 1999” ( ZSMC) . – PARALECTOTYPES (3): samedataaslectotype ( ZSMC, 3). Trogophloeus longipennis – LECTOTYPE (♀, here designated): “2006 \ longipennis \ Muséum Paris; 1906; Coll. L. Fairmaire \ Tarbes [*+43.23/+00.08*]; Pandellé; (no. 2006) \ Lectotypus; Trogophloeus; longipennis Fairm. & Lab. ; [on the back] des. Makranczy, 1999 \ ♀ \ Ochthephilus ; rosenhaueri Kiesenwetter ; det. Makranczy, 1999” ( MNHP) . – PARALECTOTYPE (1): “Tarbes; Pandellé; (no. 2006) \ remounted from; original card \ Paralectotypus; Trogophloeus; longipennis Fairm. & Lab. ; [on the back] des. Makranczy, 1999 \ ♀ \ Ochthephilus ; rosenhaueriKiesenwetter; det. Makranczy, 1999” ( MNHP, 1♀). Ancyrophorus jailensis – LECTOTYPE (heredesignated): “Iaila-Gebirge [*+44.53/+34.12*]; Krim, Moczarski \ Typus; Ancyrophorus ; jailensis; O. Scheerpeltz \ Lectotypus; Ancyrophorus ; jailensis Scheerpeltz ; [on the back] des. Makranczy, 1999 \ Ochthephilus ; rosenhaueri Kiesenwetter ; det. Makranczy, 1999”, NHMW. Ancyrophorusgracilis – HOLOTYPE: “ Carnia , Moggio, riv. Fella, 400m *+46.42/+13.20*, 23.VI.1950, leg. G. Fagel ” ( ISNB) . – PARATYPES (8): “ Carnia , Verzegnis , Intissans , rio..., 300-400m *+46.38/+12.98*, 10.VI. au 4.VII.1950, leg. G. Fagel ”, ( ISNB, 2, MNHP, 1). – “ Carnia , Venzone , torrenteVenzonazza, 300m *+46.33/+13.15*, 20.VI.1950, leg. G. Fagel ”, ( ISNB, 1). – “ Carnia , Treppo , torrente Orteglas , 1000m [*+46.53/+13.02*], 17.VI.1950, leg. G. Fagel ”, ( ISNB, 2). – “ Carnia , VillaSantina, torrente Degano, 400m [*+46.40/+12.92*], 20.VI.1950, leg. G. Fagel ” ( ISNB, 2). – “Garda-See [Lagodi Garda] *+45.67/+10.68*, leg. Breit ” ( ISNB, 1). Ancyrophoruscorsicus – HOLOTYPE: “ Corse, Vizzavona [Vizzavone], moussesderuisseaux, 900-1100m *+42.12/+09.15*, V.1950, leg. G. Fagel ” ( ISNB) . – PARATYPES (16): same data as holotype ( ISNB, 9, coll. Scheerpeltz, NHMW, 1, coll. Jarrige, MNHP, 1). – “ Corse, Vizzavona [Vizzavone], moussesdeFulminato, 900-1100m [*+42.12/+09.13*], V.1950, leg. G. Fagel ” ( ISNB, 3, coll. Scheerpeltz, NHMW, 2) .

OTHER MATERIAL: see Appendix.

REDESCRIPTION: ForebodyasinFig. 527. Measurements (n=10): HW = 0.56 (0.54-0.59); TW = 0.51 (0.49-0.53); PW = 0.59 (0.54-0.63); SW = 0.76 (0.72-0.80); AW = 0.81 (0.74-0.90); HL = 0.43 (0.40-0.45); EL = 0.20 (0.19-0.21); TL = 0.08 (0.07- 0.09); PL = 0.49 (0.46-0.52); SL = 1.07 (0.99-1.15); SC = 0.98 (0.90-1.05); FB = 2.02

(1.90-2.12); BL = 3.51 (3.20-3.79) mm. Head almost black, pronotum and abdomen blackish dark brown with occasional reddish tint. Elytra slightly reddish dark brown. Mouthparts, legs and antennae very dark brown. Body with greasy lustre mostly due to elytral setation plus forebody punctation and microsculpture. Pubescence rather fine and moderately dense, shorter and stronger (regularly spaced) on elytra, abdominal tergites with finer and longer setae, especially adjacent to laterosternites. Head anteriad eyes and near inner posterior margin of eye with stronger and darker bristles, as well as pronotal margin and middle o tibiae. Elytral apex with a few slightly longer setae near sutural corners. Last tarsomere with a few setae only.

Forebody. Antenna as in Fig. 572. Clypeus with scattered, tiny punctures (and colliculate microsculpture), trapezoid, corners rounded, anterior edge gently arched; separated by an impressed transversal line (frontoclypeal suture) across a shinier area. Supraantennal prominences well developed, feebly separated from clypeus/vertex by impressions. Vertex with oblique impressions in middle almost joining in V-shape. Temples bulging, evenly curved, barely longer than half of eye length. Neck separated by an impressed transversal groove, microsculpture much stronger than on head, with transverse cells, no setation. Pronotum with a narrow marginal bead, visible to anterior pronotal corners. Posterior pronotal angles well-formed, just slightly obtuse-angled, sides in posterior 2/3 very gently concave. 'Anchor' fully formed, longitudinal midline as a slightly elevated, impunctate, weakly microsculptured line, parallel to this line two gentle, semi-longitudinal elongate elevations in anterior half of disc. In corners of anchor feeble, oblique impressions directed outwards, in middle at sides of midline two smaller impressions. Elytra slightly broadening posteriorly, sutural corners narrowly rounded; apical sides slightly oblique and in inner halves more or less straight. Elytral surface rather even with two shallow, oblique impressions behind scutellum. Head with fine coriaceous/colliculate microsculpture, fading on elevated parts, stronger in impressions, on pronotum microsculpture slightly stronger and more even. Punctation on head rather deep but sparse, mostly confined to posterior part and sides, on pronotum also conspicuous but more evenly spaced, average interspaces 1.5x puncture diameters; elytral punctation more even and regularly spaced, average interspaces (with indistinct coriaceous microsculpture) about as puncture diameters, punctures discrete.

Abdomen. Compared to forebody, abdomen with much more sparse, finer, less distinct punctation, microsculpture on tergal apices fine coriaceous with moderately transverse cells. Tergite VII posterior margin with palisade fringe unmodified in middle (nearly uniform breadth). Tergite VIII (Fig. 231) basal edge evenly arched, but straight at a short distance in middle of basal sclerotized band; apical edge with sinuate (protruding) corners, and broad, moderately deep emargination in between. Sternite VIII with rounded apical corners, apex in males shallowly concave laterally, gently sinuate in middle; in females slightly more sinuate (convex) in middle. Tergite X unmodified, apex very slightly wider in males than in females. Aedeagus as in Fig. 212, inner sclerites as in Figs 213-214. Female ringstructures as in Figs 215-216.

COMPARATIVE NOTES: This species is extremely similar to O. praepositus , and although their distribution ranges do not overlap perfectly, they are often found in the same microhabitats and can be separated with most difficulty. Apart from the more slender antennae and more elongate elytra of O. rosenhaueri , in doubtful situations males are best separated by the inner sclerites of the aedeagi, females by the slightly different form of tergites IX. Other possibly confused species (by similar size and slender antenna) is O. lenkoranus , but it is very different in the inner sclerite of the aedeagus and the female ringstructure, in most specimens by the basal margin of tergite VIII. The specimens known from Corse ( Corsica) have much more slender/elongate antennae, but male genitalia are identical to those in the rest of the distribution range.

NOTE: Dr. D. Drugmand (Brussels) kindly provided photocopies of the relevant pages from L. Pandellé's locality key (a handwritten notebook), which allowed adequate labeling of the types of Trogophloeus longipennis .

DISTRIBUTION: Known from France (Alpes-Maritimes and Corsica) through the Alps and northern Carpathians to the whole northeastern Mediterranean Basin (including Sicily) to the Caucasus.

BIONOMICS: Specimens were collected from willow moss (Fontinalis sp.) on streambank, entirely soaked moss, moss at waterfall, wet sand and gravel, flood debris, muddy spots on bank of small stream in mixed forest, under stones on riverbank.

Ochthephilus solodovnikovi Gildenkov, 2000 Figs 12, 159-163, 175-176, 535, 584 Ochthephilus solodovnikovi Gildenkov, 2000: 1185 . – Gildenkov, 2000b: 847.

TYPE MATERIAL EXAMINED: Ochthephilus solodovnikovi – PARATYPES (2): “Краснодарск.[ий] кр.[ай] Красная Поляна р.[ека] Галясы, 1400м, 23.VIII.1995 М. Савицкий,” (coll. Solodovnikov, ZMUC, 13, 1♀) .

OTHER MATERIAL: RUSSIA: Краснодарский край, 7км NW Бабук-Аула, Ю. скл. г. Хуко [7km NW Babuk-Aul, S slopes of Mt. Khuko], 1600м *+43.94/+39.79*, 24.VI.1994, leg. А. Солодовников, forest, stream edge, coll. Solodovnikov, ZMUC (13, 1). – NW Caucasus, 12km E of vill. Krasnaya Polyana, forest zone, 700-800m *+43.66/+40.35*, 29.VII.1994, leg. A. Solodovnikov, coll. Solodovnikov, ZMUC (1). – W Caucasus, vill. Temnoleskaya, 45km S of Maykop, 700m *+44.21/+40.00*, 8.VI.1999, leg. A. Solodovnikov coll. Solodovnikov, ZMUC (1), FMNH (13), MHNG (13, 1♀) , NHMW (1), HNHM (13, 1♀, 1), coll. Schülke (13). — GEORGIA: Abkhazia , Gumistinskiy Nature Reserve , Tsumuri, Vostochnaya Gumista River bank, 420m *+43.23/+41.07*, 22.VII.1990, leg. V.I. Gusarov, coll. Gusarov, ZMUN (2). – Cauc. min. bor. Trialetskij Chreb., Bakuriani, 1800-2200m *+41.71/+43.54*, 4-7.VII.1986, leg. D.W. Wrase & M. Schülke, coll. Schülke (2♀) , HNHM (13).

REDESCRIPTION: Forebody as in Fig. 535. Measurements (n=10): HW = 0.67 (0.64-0.69); TW = 0.62 (0.59-0.665); PW = 0.81 (0.78-0.85); SW = 0.90 (0.84-0.95); AW = 1.03 (0.98-1.13); HL = 0.50 (0.48-0.51); EL = 0.21 (0.19-0.22); TL = 0.12 (0.11- 0.13); PL = 0.62 (0.58-0.65); SL = 1.09 (1.02-1.15); SC = 0.99 (0.93-1.06); FB = 2.29 (2.12-2.39); BL = 4.18 (3.62-4.51) mm. Body uniformly very dark brown, almost black, but with a distinct reddish tint. Rare lighter specimens with only basal antennomeres, mouthparts and legs reddish medium to dark brown, elytra occasionally more reddish dark brown. Body with slightly above average lustre mostly due to occasional shinier elevated spots on head and pronotum. Pubescence rather fine and moderately dense, shorter and stronger (regularly spaced) on elytra, abdominal tergites with finer and longer setae, especially adjacent to laterosternites. Head anteriad eyes and near inner posterior margin of eye with stronger and darker bristles, as well as pronotal margin and middle of tibiae. Elytral apex with a couple of very long setae near sutural corners. Last tarsomere with a few setae only.

FIGS 281-285

TergitesX; Ochthephilusangustior (Bernhauer), 3 (281), same, ♀ (282), O. omalinus (Erichson) , 3 (283), same, ♀ (284), O. szeli sp. n., 3 (285). Scalebar = 0.1 mmfor 281-282, 0.115 mmfor 283-285.

Forebody. Antenna as in Fig. 584. Clypeus sparsely and finely punctate (colliculate microsculptured), trapezoid, corners rounded, anterior edge gently arched; separated by an impressed transversal line (frontoclypeal suture) across a shinier area. Supraantennal prominences well developed, feebly separated from clypeus/vertex by impressions. Vertex with a few isolated impressed spots. Temples bulging, evenly curved, noticeably longer than half of eye length. Neck separated by an impressed transversal groove, microsculpture much stronger than on head, with transverse cells, no setation. Pronotum with a narrow marginal bead, visible to anterior pronotal corners. Posterior pronotal angles well-formed, just slightly obtuse-angled, sides in posterior 2/3 gently concave/bisinuate. 'Anchor' fully formed, longitudinal midline as a slightly elevated, impunctate, weakly microsculptured line, parallel to this line two gentle, semi-longitudinal elongate elevations in anterior half of disc. In corners of anchor feeble, oblique impressions directed outwards, in middle at sides of midline two smaller impressions. Elytra slightly broadening posteriorly, sutural corners narrowly rounded; apical sides slightly oblique and in inner halves more or less straight. Elytral with two shallow, oval, oblique impressions behind scutellum and disc shallowly, longitudinally impressed, connecting larger, slightly impressed anterior and posterior halves (behind border of latter punctation abruptly changes to much finer). Head with fine coriaceous/colliculate microsculpture, fading on elevated parts, stronger in impressions, on pronotum microsculpture slightly stronger and more even. Punctation on head deep but sparse, on pronotum more evenly spaced, average interspaces much larger than puncture diameters; elytral punctation more even and regularly spaced, average interspaces (with indistinct coriaceous microsculpture) about as puncture diameters, punctures discrete.

Abdomen. Compared to forebody, abdomen with much more sparse, finer, less distinct punctation, microsculpture on tergal apices fine coriaceous with moderately transverse cells. Tergite VII posterior margin with palisade fringe unmodified in middle (nearly uniform breadth). Tergite VIII basal edge in females (Fig. 176) more strongly angular, slightly more evenly arched in males (Fig. 175); apical edge with sinuate (protruding) corners, and broad, moderately deep emargination in between. Sternite VIII with rounded apical corners, apex in males shallowly concave laterally, gently sinuate in middle; in females slightly more sinuate (convex) in middle. Tergite X unmodified, apex very slightly wider in males than in females. Aedeagus as in Fig. 159, inner sclerites as in Figs 160-161. Female ringstructures as in Figs 162-163.

COMPARATIVE NOTES: A very characteristic, large sized species, easily distinguishable by the wide pronotum with conspicuous marginal bead and most importantly the basal formation of tergite VIII that only group it with the much smaller O. lenkoranus .

DISTRIBUTION: The species is so far known from the greater area of the

Caucasus.

BIONOMICS: Specimens were collected at forest stream, on stream- and riverbank.

Ochthephilus tatricus ( Smetana, 1967) Figs 184-188, 202, 533, 562 Ancyrophorus tatricus Smetana, 1967: 384 , 386.

Ochthephilus tatricus (Smetana) . – Lohse & Lucht, 1989: 400. – Makranczy, 2001: 179.

TYPEMATERIALEXAMINED: HOLOTYPE (♀): “Slov.[akia] Vys.[oké] Tatry, Tatr.[anská] Lomnica [ca. 900m]; Smetana 1.VII.1956 a” (coll. Smetana, CNCI, no. 16784). – PARATYPE (1): same data as holotype (coll. Smetana, CNCI, 13) .

OTHERMATERIAL: FRANCE: Savoie, Plan-Lachat, s/ Valloire , 2000m *+45.09/+06.43*, 29.VIII.1967, leg. A. Comellini, MHNG (13) . – dept. Alpes-Maritimes, Roquebillière (c6), valléeGordolasque *+44.04/+07.36 *, 21.V.1969, leg. MTronquet, détrit. inond., coll. Tronquet (13). — GERMANY: Schwarzwald, Bonndorf *+47.82/+08.34 *, VI.1912, leg. W. Sattler, coll. Sattler, SMFD (13) . — SWITZERLAND: Kt . Wallis, Walliser Alpen, Zinal , 1700m *+46.14/+07.62 *, 5.VII.2002, leg. Mehnert, Nadelwald, Bachufer, inMoos, coll. Gollkowski (1). — AUSTRIA : Steiermark, Hochschwabgebiet, Unt. Ring, 960m, Gesiebe hohler Ahorn *+47.63/+15.18*, 3.VII.1992, leg. A. Kapp, coll. Kapp (1) . – Tirol, Stanzach / Lech, Namloser Tal *+47.37/+10.62*, 24.V.1995, leg. K. Renner, Autokätscher, coll. Gollkowski (1), coll. Renner (1), HNHM (1) . – Tyr., Kitzbühel *+47.45/+12.39*, 24.V.1995, leg. Breit, coll. Scheerpeltz, NHMW (13) . – Kärnten, GailtalerAlpen *+46.63/+13.56*, coll. Bierig, FMNH (1) . — ITALY: Trentino , ETrento, NRoncegno, 700-1600m *+46.08/+11.39 *, 26.VI.1995, leg. V. Assing, Autokescher, coll. Assing (13). – Vestenavecchia / Tregnago ( VR) (car net) *+45.54/+11.19 *, 20.IV.1996, leg. A. Zanetti, coll. Zanetti (1). – Dolomiti ( TN), SanMartinodi Castrozza *+46.63/+13.56 *, 29-31.VII.1980, leg. F. Montemurro (mt 1467), HNHM (13) . — SLOVENIA: SteinerAlpen, LogarskaDolina , 810m *+46.40/+14.63 *, 5.VI.1999, leg. A. Kapp, coll. Kapp (1). — BOSNIA-HERZEGOVINA: Cernesnica pl., [pečina] Ledenica *+44.61/+16.44*, NHMW (1♀). – Bosnien, KraljevskaSutjesca-Zadolci *+44.12/+18.20 *, 5.V.1990, leg. P. Wunderle, autokätscher, coll. Wunderle (13), SDEI (1♀). — SLOVAKIA: Branisko, Sl'ubica, 600-900m *+48.99/+20.87 *, 16.VI.1984, leg. J. Janák, coll. Janák (1). – ZávadkanadHronomenv., Machnatádolina *+48.79/+19.97*, 24.V.2009, leg. M. Mantič, moss on stones in brook, coll. Mantič (1). — ROMANIA: jud . Braşov, Culoarul Rucǎr-Bran, 2.0 km SWFundǎţica, PeşteraUlucelor , streamfromentrance, 45°24'35"N, 25°15'47"E, 950 m, 16.VII.2011, leg. Gy. Makranczy (459), moss and leaflitter at rocks in small cascades, flotation, HNHM (1) . — UKRAINE: Carpath. or., Worochta [Vorokhta], Prut [river] *+48.29/+24.57*, 12.VI.1911, leg. Lokay, coll. Lokay, NMPC (1).

REDESCRIPTION: ForebodyasinFig. 533. Measurements (n=10): HW = 0.59 (0.54-0.60); TW = 0.55 (0.51-0.56); PW = 0.66 (0.58-0.69); SW = 0.76 (0.72-0.79); AW = 0.86 (0.83-0.90); HL = 0.44 (0.40-0.48); EL = 0.15 (0.14-0.18); TL = 0.12 (0.10- 0.14); PL = 0.53 (0.48-0.57); SL = 0.96 (0.90-1.03); SC = 0.88 (0.82-0.94); FB = 2.01 (1.88-2.10); BL = 3.48 (3.00-4.20) mm. Head, pronotumandabdomendarkbrown with slight reddish tint; head often darker, almost black. Elytra somewhat reddish medium to dark brown, same colour as antennae, mouthparts and legs; in case of lighter specimens scutellar area to shoulders darker and epistomal sulcus appears as transversal black line. Body appearing rather dull, especially head and pronotum with fine but very strong microsculpture. Pubescence extremely fine and moderately dense, shorter and stronger (regularly spaced) on elytra, abdominal tergites with fine and much longer setae on apices. Head anteriad eyes and near inner posterior margin of eye with stronger and darker bristles, as well as pronotal margin and middle of tibiae. Elytral apex with at least two conspicuously longer setae near sutural corners. Last tarsomere with a few setae only.

Forebody. Antenna as in Fig. 562. Clypeus almost impunctate (colliculate microsculptured), trapezoid, corners rounded, anterior edge gently arched; separated by impressed transversal line (frontoclypeal suture) across a shinier area. Supraantennal prominences well developed, feebly separated from clypeus/vertex by impressions. Vertex with oblique impressions in middle almost joining in U-shape. Temples slightly bulging, evenly curved, little longer than half of eye length. Neck shiny, separated only by a weakly impressed transversal groove, no setation. Pronotum with a narrow marginal bead, visible to anterior pronotal corners. Posterior pronotal angles well-formed, just slightly obtuse-angled, sides in posterior 2/3 very gently concave/bisinuate. 'Anchor' fully formed, longitudinal midline as a slightly elevated, impunctate, weakly microsculptured line, parallel to this line two gentle, semi-longitudinal elongate elevations in anterior half of disc. In corners of anchor feeble, oblique impressions directed outwards, in middle at sides of midline two smaller impressions. Elytra slightly broadening posteriorly, sutural corners narrowly rounded; apical sides slightly oblique and in inner halves more or less straight. Elytral surface rather even with two shallow, very elongate impressions behind scutellum. Head with fine coriaceous/colliculate microsculpture, fading on elevated parts, stronger in impressions, on pronotum microsculpture slightly stronger and more even. Punctation on head shallow and sparse, on pronotum more evenly spaced, average interspaces much larger than puncture diameters; elytral punctation much deeper, more even and regularly spaced, average interspaces (with indistinct coriaceous microsculpture) about as puncture diameters, punctures discrete.

Abdomen. Compared to forebody, abdomen with much more sparse, finer, less distinct punctation, microsculpture on tergal apices fine coriaceous with moderately transverse cells. Tergite VII posterior margin with palisade fringe unmodified in middle (nearly uniform breadth). Tergite VIII (Fig. 202) basal edge evenly arched, with small concavity in middle of basal sclerotized band; apical edge with sinuate (protruding) corners, and broad, moderately deep emargination in between. Sternite VIII with rounded apical corners, apex in males shallowly concave laterally, gently sinuate in middle; in females slightly more sinuate (convex) in middle. Tergite X unmodified, apex very slightly wider in males than in females. Aedeagus as in Fig. 184, inner sclerites as in Figs 185-186. Female ringstructures as in Figs 187-188.

COMPARATIVE NOTES: This species is similar to O. brachypterus and O. legrosi in having the very dull pronotum with obscured punctation, but differs from them by the elongate (narrower) and conspicuously parallel-sided (straight) elytra as well as usually being the smallest of the three.

DISTRIBUTION: Known from France (Alpes-Maritimes) through the Alps, northern Italy, the Carpathians to the northern Balkans.

BIONOMICS: This very rare species (almost always collected as singleton) lives in wet moss at streams, together with the common O. praepositus . The habitat is not confined to higher elevation (like in the related O. brachypterus ) but bound to caves or shaded environments of abundant underground cavities. On rare occasions specimens were captured in flight. The collector of the holotype gave the following information (A. Smetana, pers. comm.): the type specimens were collected in one of the shady, small valleys in coniferous forest just above Tatranská Lomnica. He found it in entirely water soaked wet moss on big rocks in wild creeks. This confirms my observation that this species indeed lives at cold, humid, forest-covered, densely vegetated habitats; not only in moss, but wet leaflitter. A great percentage of the known specimens was collected by car-nets.

FIGS 286-299

(286-290) Ochthephilusgusarovi sp. n.; aedeagus (286), innersclerites (287-288), femaleringstructures (289-290). (291-295) O. venustulus (Rosenhauer) ; aedeagus (291), innersclerites (292-293), femaleringstructures (294-295). (296-299) O. qingyianus sp. n.; aedeagus (296), innersclerite (297), femaleringstructures (298-299). Scalebar = 0.05 mmfor 298-299, 0,055 mmfor 289-290, 0.06 mmfor 294-295, 0.1 mmfor 286-288, 296-297, 291-293.