Aulacus insularis, Jennings & Parslow & Austin, 2018

Jennings, John T., Parslow, Ben A. & Austin, Andrew D., 2018, Systematics of the parasitoid wasp genus Aulacus Jurine (Hymenoptera: Evanioidea: Aulacidae) from Australia, Zootaxa 4538 (1), pp. 1-113 : 7-10

Key to Australian Aulacus Jurine (females only)

We have not included in the key four species known only from males, viz., A. flindersbaudini Jennings, Austin & Stevens , a largely reddish brown species which is endemic to Kangaroo Island, South Australia; A. insularis sp. nov., which is readily separated from all other Australian species by the two distinctive cream bands on metasomal segments 2 and 3 ( Fig. 26A ); A. naumanni sp. nov., which has a colour combination different from all other Australian species (see comments below under this species), and A. vespiformis Kieffer , which has distinctive banding on the metathorax ( Figs 50A, B ) and anterior half of fore wing distinctively fuscous ( Fig. 50A ).

1 Mesoscutum in lateral view angular antero-dorsally (e.g., Fig. 18E ).......................................... 2

- Mesoscutum in lateral view rounded antero-dorsally (e.g., Fig. 2E ).......................................... 20

2(1) Metasoma clavate ( Figs 9A, B ); face punctate-reticulate; fore wing vein 2-Rs+M very long; mesothorax pale orange in the anterior half and largely dark brown in the posterior half; brown band either side of fore wing veins 1-Rs and 1-M and apically in costal cell [NSW and Queensland]................................................ A. bicolor sp. nov.

- Metasoma ovate (e.g., Figs 29A, B ); other features not in combination........................................ 3

3(2) Fore wing with brown infuscation (spot) at apex (e.g., Fig. 1 )............................................... 4

- Fore wing lacking brown infuscation (spot) at apex...................................................... 15

4(3) Hind wing venation incomplete, variable loss of one or more veins M+Cu, Cu, r-m and 2-M....................... 5

- Hind wing venation complete (e.g., Fig. 1 ).............................................................. 7

5(4) Hind tarsal claw long, 3.0× length tarsomere 5; face rugose with a few punctures medially; head and mesosoma black, metasoma orange; ovipositor guide about 2/3 rd distance along coxa, oblique ( Fig. 51F ) [Victoria]..... A. walkeri sp. nov.

- Hind tarsal claw short, about half length of tarsomere 5; other features not in combination......................... 6

6(5) Face rugose; head and mesosoma blackish brown, metasomal T1 and T2 orange, remainder of metasoma blackish brown ( Figs 29A, B ); ovipositor guide about ¾ way along coxae, oblique, distal fringe of long setae, long setae in groove [NSW]............................................................................ A. kiwarrakensis sp. nov.

- Face punctate-rugose; head and mesosoma black, metasoma orange ( Figs 30A, B ); ovipositor guide medial, oblique, setae on lower distal margin, lacking setae in groove [Tasmania]...................................... A. leai sp. nov.

7(4) Ovipositor guide on hind coxa transverse (e.g., Fig. 5D ).................................................... 8

- Ovipositor guide on hind coxa oblique (e.g., Fig. 51F )..................................................... 9

8(7) Ovipositor guide on hind coxa distal; face rugulose-reticulate; frons without lateral medial carina above toruli; head black, meso- and metasoma orange ( Figs 36A, B ) [Victoria]....................................... A. neboissi sp. nov.

- Ovipositor guide on hind coxa medial; face punctate-rugose; frons with lateral medial carina above toruli; largely dark black-brown species ( Figs 25A, B ) [Queensland]........................................... A. hackeri sp. nov.

9(7) Ovipositor guide on hind coxa distal (e.g., Fig. 12F )...................................................... 10

- Ovipositor guide on hind coxa medial (e.g., Fig. 7F )...................................................... 14

10(9) Frons with lateral medial carina above toruli; face imbricate; head, scape, pedicel, propleuron, legs and metasoma orange, except for variable amounts of black on metasomal T1, mesosoma black ( Figs 18A, B ) [NSW and Victoria]............................................................................................. A. elegans sp. nov.

- Frons without lateral medial carina above toruli; other features not in combination.............................. 11

11(10) Face imbricate; vertex punctate ( Fig. 33C ); posterior margin of head distinctly concave in dorsal view ( Fig. 33C ); metasoma dark brown, head, scape, pedicel, flagellomere 1 and propodeum reddish brown, legs and metasoma brown ( Figs 33A, B ) [NSW and Queensland]....................................................... A. minutus sp. nov.

- Face other than imbricate; other features not in combination............................................... 12

12(11) Face rugulose; body length 8.1 mm, excluding ovipositor; ovipositor 5.0 mm; head and mesosoma black, metasoma largely orange, ovipositor sheaths black, ovipositor brown ( Figs 45A, B ) [Tasmania]................ A. smithi sp. nov.

- Face rugulose-reticulate or rugose-reticulate; other features not in combination................................ 13

13(11) Body length excluding ovipositor, 8.0 mm; ovipositor 5.3 (5.0–5.7) mm; vertex rugose-reticulate, almost strigate-reticulate ( Fig. 53C ); head and mesosoma black, metasomal T1 and T2 largely orange with remainin g segments blackish brown ( Figs 53A, B ) [NSW]............................................................... A. williamsi sp. nov.

- Body length excluding ovipositor, 6.5 mm; ovipositor 3.7 mm; vertex rugose-reticulate, punctate-reticulate near ocelli ( Fig. 35C ); body orange-brown [Queensland]................................................ A. nebo sp. nov.

14(9) Body length, excluding ovipositor, 9.45 (9.4–9.5) mm; ovipositor longer than body, 12.5 (12.0–13.0) mm [Victoria]...................................................................................... A. confusus sp. nov.

- Body length, excluding ovipositor, 7.5 mm; ovipositor shorter than body, 5.7 mm [Queensland].................................................................................................. A. mareebaensis sp. nov.

15(3) Face with punctures, sculpturing either punctate-rugose or punctate-reticulate................................. 16

- Face lacking punctures, sculpturing either rugose or rugulose.............................................. 17

16(15) Face punctate-rugose; antenna black except flagellomeres 4-7 cream ( Figs. 43A, B ), ovipositor sheath brown; metasoma 1.3× length of mesosoma, and T1 and T2 narrow, glabrous, smooth tending to imbricate with scattered, small punctures posteriorly [Queensland].............................................................. A. scitulus sp. nov.

- Face punctate-reticulate; antenna black except flagellomeres 5–7 white ( Figs 37A, B ); ovipositor sheath black except tip pale orange; metasoma 1.19 (1.12–1.21)× length of mesosoma, and T1 and T2 broad, several shallow and indistinct punctures antero-dorsally Tasmania, Victoria, Queensland]................................... A. obcordellus sp. nov.

17(15) Face rugulose (wrinkles fine); head, meso- and metasoma black, tibae and tarsi largely light brown; body length, excluding ovipositor, 10.5 mm [South Australia]................................................. A. moerens Westwood

- Face rugose (wrinkles coarse, e.g., Fig. 27C ); body length excluding ovipositor, ≤ 9.0 mm....................... 18

18(17) Metasomal T1 and T2 rugulose dorsally, smooth laterally, lacking punctures; body length 7.6 mm, excluding ovipositor [ovipositor and sheath largely missing on holotype]; black except antenna orange-yellow, mandibles, fore and mid trochanters and femora brown, fore and mid tibiae and tarsi pale brown, hind legs dark brown, metasoma dark brown ( Fig. 21A ) [Victoria]................................................................. A. flavimanus (Kieffer)

- Metasomal T1 and T2 with at least a few punctures; other features not in combination........................... 19

19(18) Metasomal T1 and T2 glabrous, except for a few indistinct, shallow punctures; body length 6.8 (5.8–7.3) mm, excluding ovipositor; ovipositor 2.8 (2.30–3.35) mm; ovipositor guide absent [Victoria]..................... A. brabyi sp. nov. - Metasomal T1 and T2 weakly rugulose-punctate dorsally; body length 9.0 mm, excluding ovipositor; ovipositor short, 3.5 mm; ovipositor guide medial, transverse [Victoria]........................................ A. flavicornis Kieffer

20(1) Metasoma clavate (e.g., Figs 9A, B ).................................................................. 21

- Metasoma ovate (e.g., Figs 29A, B )................................................................... 38

21(20) Fore wing with brown spot at apex (see Fig. 1 )........................................................ 23

- Fore wing without brown spot at apex................................................................. 33

22(21) Hind wing venation incomplete, veins M+Cu, Cu, r-m and 2-M absent....................................... 23

- Hind wing venation complete, veins M+Cu, Cu, r-m and 2-M present (e.g., Fig. 1 ).............................. 24

23(22) Ovipositor guide absent ( Fig. 15F ); ovipositor 6.3 (6.0–6.5) mm; metasoma 1.80 (1.60–1.92)× length of mesosoma; metasoma dark brown [Victoria]..................................................... A. dandenongensis sp. nov.

- Ovipositor guide medial, oblique; ovipositor very short, 3.95 mm; metasoma 1.5× length of mesosoma; metasoma dark blackish brown with pale brown band dorsally before posterior margin of T2 ( Fig. 28B ) [Queensland].................................................................................................. A. kittelae sp. nov.

24(22) Ovipositor guide absent; posterior margin of head not concave in dorsal view ( Fig. 54C ) [Queensland]................................................................................................ A. wrightae sp. nov.

- Ovipositor guide present, either transverse or oblique; posterior margin of head concave in dorsal view............. 25

25(24) Ovipositor guide transverse......................................................................... 26

- Ovipositor guide oblique........................................................................... 28

26(24) Head with lateral medial carina present above toruli (e.g., Figs 11C , 43F )..................................... 27

- Head without lateral medial carina above toruli [South Australia]........... A. belairensis Jennings, Austin & Stevens

27(25) Ovipositor guide somewhat caudad, narrow ( Fig. 31G ); head and mesosoma black, metasoma orange ( Figs 31A, B ) [Queensland, NSW, ACT, Victoria]................................................. A. longiventris (Kieffer)

- Ovipositor guide medial, broad; body orange-brown [WA]................... A. houstoni Jennings, Austin & Stevens

28(25) Occipital carina present laterally, weak (narrow) ( Fig. 7D ); body length excluding ovipositor 7.5 mm; ovipositor short, 3.0 mm [Queensland]................................................................ A. bamagensis sp. nov.

- Occipital carina absent; other features not in combination................................................. 29

29(28) Face with punctures............................................................................... 30

- Face without punctures............................................................................. 31

30(28) Face punctate-reticulate; frons without lateral medial carina above toruli; metasomal T2 with distinctive cream band dorso-laterally ( Figs 17A, B ) [Queensland]................................................. A. doddi sp. nov.

- Face punctate-rugose; frons with lateral medial carina above toruli; body yellowish-brown, including metasomal T2 ( Figs 41A, B ) [Queensland]................................................................. A. quickei sp. nov.

31(29) Head and mesosoma red-brown ( Figs 27A, B ); face rugose ( Fig. 27C ); ovipositor guide somewhat medial [NSW]...................................................................................... A. jamberoo sp. nov.

- Head and mesosoma largely black; other features not in combination........................................ 32

32(31) Body black except femora, tibiae and posterior metasomal T1 orange, tarsomeres predominantly white ( Figs 2A, B ); face rugose; ovipositor guide somewhat caudal [Queensland and NSW].......................... A. albimanus (Kieffer)

- Body black except propleuron and ventral half of pronotum orange, metasomal T1 and T2 orange, and remainder dark brown ( Figs 13A, B ); face rugulose; ovipositor guide medial [Queensland]........................ A. burnsi sp. nov.

33(21) Hind wing venation incomplete...................................................................... 34

- Hind wing venation complete (see Fig. 1 ).............................................................. 35

34(33) Ovipositor guide transverse; head and mesosoma dark brown, metasoma yellow brown [Western Australia]............................................................................ A. mcmillani Jennings, Austin & Stevens

- Ovipositor guide somewhat oblique; head and mesosoma black, metasoma orange ( Figs 24A, B ) [Queensland]........................................................................................... A glorious sp. nov.

35(33) Occipital carina present laterally, weak (narrow); face punctate-rugulose; mesosoma and metasoma about equal in length; ovipositor guide more or less oblique [South Australia]....................... A. grossi Jennings, Austin & Stevens

- Occipital carina absent; other features not in combination................................................. 36

36(35) Frons with lateral medial carina above toruli [Tasmania]..................................... A. simsoni sp. nov.

- Frons without lateral medial carina above toruli......................................................... 37

37(36) Face punctate-rugose ( Fig. 42C ); body dark brown-black [ACT, NSW, Victoria].................... A. rieki sp. nov.

- Face without punctures, rugose-reticulate; head and mesosoma black, metasoma orange except for variable amounts of black on T1 ( Figs 23A, B ) [NSW, Queensland]........................................ A. fuscicornis (Cameron)

38(21) Ovipositor guide absent............................................................................ 39

- Ovipositor guide present............................................................................ 42

39(38) Frons with lateral carina above toruli (e.g., Figs 11C , 43F )................................................. 40

- Frons without lateral carina above toruli............................................................... 41

40(39) Face rugulose-reticulate; head and mesothorax black, metasoma dark brown ( Figs 38A, B ) [NSW].............................................................................................. A. pallidicaudis (Cameron)

- Face rugose; body black except legs, and metasomal T1 and T2 with variable amounts of light brown ( Figs 4A, B ) [Queensland, South Australia]................................................................ A. aquilus sp. nov.

41(39) Face punctate-reticulate; head and mesosoma brown except some orange on gena, propleuron and lateral pronotum, scape, pedicel, legs and metasoma orange ( Figs 22A, B ) [NSW].................................... A. froggatti sp. nov.

- Face rugose; body black except scape orange, fore and mid trochanters dark orange, all femora orange, fore and mid tar- someres 1–4 cream, all tarsomeres 5 brown, hind tibiae dark brown and first two-thirds basitarsus dark brown, apical onethird basitarsus and tarsomeres 2–4 cream ( Figs 8A, B ) [Tasmania]........................... A. bashfordi sp. nov.

42(38) Ovipositor guide transverse......................................................................... 43

- Ovipositor guide oblique........................................................................... 45

43(42) Body entirely black ( Fig. 42A ); length 11.4 mm, excluding ovipositor; ovipositor 17.0 mm [NSW, Queensland]..................................................................................... A. planiceps (Szépligeti)

- Body generally dark brown or head largely orange and mesosoma black...................................... 44

44(43) Length 6.2 (5.6–6.6) mm, excluding ovipositor; ovipositor 7.6 (7.5–7.7) mm; head orange except for variable amounts of dark brown medially on frons and vertex, mesosoma black ( Figs 16A, B ) [Victoria]................ A. deansi sp. nov.

- Length 12.5 mm, excluding ovipositor; ovipositor long, 14.3 mm; generally dark brown, with variable amounts of black on frons and face, pronotum, mesoscutum, metanotum, mesopleuron, mesepimeron, basally on mid and hind coxae, hind trochanter and trochantellus ( Figs 19A, B ) [Queensland]...................................... A. festivus (Kieffer)

45(42) Ovipositor guide somewhat distal.................................................................... 46

- Ovipositor guide medial............................................................................ 48

46(45) Face rugose, punctate-rugose laterally; ovipositor guide distal ( Fig. 12F ); length 7.3 mm, excluding ovipositor; ovipositor 9.7 mm; head and metasoma dark brown, mesosoma black [ACT].............................. A. broadi sp. nov.

- Face lacking punctures; other features not in combination................................................. 47

47(46) Face rugulose-reticulate; head and mesosoma black, metasoma black except somites 1–3 orange, with variable amounts of black anteriorly on T1 ( Figs 10A, B ); metasoma 1.22 (1.15–1.26)× length of mesosoma [NSW]................................................................................................ A. boonanghiensis sp. nov.

- Face rugulose; head black, meso- and metasoma orange ( Figs 6A, B ); metasoma 1.3 (1.2–1.4)× length of mesosoma [Tasmania]............................................................................. A. atriceps Kieffer

48(45) Head black...................................................................................... 49

- Head orange to dark brown......................................................................... 54

49(48) Mesosoma and metasoma orange..................................................................... 50

- Mesosoma black, metasoma variable in colour, but not entirely orange....................................... 51

50(49) Face rugulose; length 9.6 mm, excluding ovipositor; ovipositor long, 10.6 mm [Queensland]...... A. umbackae sp. nov.

- Face punctate-rugose; length 8.0 mm, excluding ovipositor; ovipositor short, 6.5 mm [Tasmania]................................................................................................... A. warraensis sp. nov.

51(49) Ovipositor short, <6.0 mm.......................................................................... 52

- Ovipositor long,> 9.0 mm.......................................................................... 53

52(51) Length 7.0 mm, excluding ovipositor; ovipositor short, 5.5 mm; head 1.35× wider than long when viewed dorsally; face rugulose; metasoma predominantly dark brown, T1 and T2 paler ( Fig. 48A ) [Victoria].......... A. truncatus (Kieffer)

- Length 7.1 mm, excluding ovipositor; ovipositor 5.5 mm; head 1.10× wider than long when viewed dorsally; face rugose; metasoma dark brown ( Fig. 46B ) [Tasmania]........................................... A. tasmanicus sp. nov.

53(51) Metasoma 1.6 (1.5–1.7)× length of mesosoma; ovipositor 11.0 (10.5–11.5) mm; metasoma with T1 black, T2 orange, T3– T8 dark brown ( Fig. 43B ) [ACT and NSW]................................................. A. anici sp. nov.

- Metasoma 1.3× length of mesosoma; ovipositor 9.4 mm; metasoma dark brown except T2 generally paler ( Fig. 5B ) [Queensland]....................................................................... A. aroueti (Girault)

54(48) Large specimens, length 16.5 mm, excluding ovipositor; ovipositor 18.8 mm; head and body largely orange-red; ovipositor guide medial and oblique [Western Australia]............................. A. douglasi Jennings, Austin & Stevens

- Small specimens. length, excluding ovipositor, less than 10 mm; other features not in combination................. 55

55(54) Head dark brown, meso- and metasoma pale orange ( Figs 32A, B ); length 6.8 mm, excluding ovipositor; ovipositor 6.1 mm; metasoma 1.4× length of mesosoma [Victoria]........................................ A. pallidus sp. nov.

- Head and metasoma orange, mesosoma brown ( Figs 47A, B ); length 7.6 (6.5–8.25) mm, excluding ovipositor; ovipositor 13.6 (13–14.2) mm; metasoma 1.5× length of mesosoma [Victoria]............................. A. tiernyi sp. nov.

Figures

FIGURE 1. Wings of Aulacus neboissi sp. nov. showing complete hind wing venation. Scale bar = 1 mm.

FIGURE 2. Aulacus albimanus (Kieffer), holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) lateral mesosoma; (F) inner side right hind coxa.

FIGURE 4. Aulacus aquilus sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) lateral mesoscutellum; (F) dorsal hind coxae.

FIGURE 5. Aulacus aroueti (Girault), holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorso-lateral metasoma; (D) inner side of right coxa.

FIGURE 6. Aulacus atriceps Kieffer, holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) lateral mesosoma; (F) dorsal hind coxae.

FIGURE 7. Aulacus bamagensis sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus. Aulacus bamagensis sp. nov., paratype, (C) dorsal head; (D) postero-lateral head; (E) dorsal mesosoma; (F) dorsal hind coxae.

FIGURE 8. Aulacus bashfordi sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) lateral head and mesosoma; (D) dorsal hind coxae.

FIGURE 9. Aulacus bicolor sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) dorsal mesosoma and metasomal T1 and T2; (E) lateral mesosoma; (F) inner lateral right hind coxa.

FIGURE 10. Aulacus boonanghiensis sp. nov., paratype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) dorsal mesosoma; (E) lateral head and mesosoma; (F) dorso-lateral hind coxa.

FIGURE 11. Aulacus brabyi sp. nov., paratype female, (A) dorsal habitus; (B) lateral habitus; (C) fronto-dorsal head; (D) lateral head; (E) dorso-lateral mesosoma; (F) dorsal hind coxae.

FIGURE 12. Aulacus broadi sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head and mesosoma; (E) dorsal mesosoma; (F) dorsal hind coxae.

FIGURE 13. Aulacus burnsi sp. nov., paratype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) dorsal mesosoma, holotype, (F) dorsal hind coxae.

FIGURE 15. Aulacus dandenongensis sp. nov., paratype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head and mesosoma; (E) dorsal propleuron and mesosoma. Aulacus dandenongensis sp. nov., holotype female, (F) inner left hind coxa.

FIGURE 16. Aulacus deansi sp. nov., paratype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) dorsal mesosoma; (F) dorsal hind coxae.

FIGURE 17. Aulacus doddi sp. nov., paratype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head and mesosoma; (E) dorsal mesosoma; (F) dorsal hind coxae.

FIGURE 18. Aulacus elegans (Kieffer), paratype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) lateral pronotum and mesoscutum; (F) dorsal hind coxae.

FIGURE 19. Aulacus festivus (Kieffer), female, (A) dorsal habitus; (B) lateral habitus; (C) frontal head; (D) lateral head; (E) dorsal mesosoma; (F) lateral mesosoma; (G) dorsal hind coxae; (H) dorsal metasomal T1.

FIGURE 21. Aulacus flavimanus (Kieffer), holotype female, (A) lateral habitus; (B) dorsal habitus; (C) lateral metasoma (Images ©The Trustees of the Natural History Museum, London).

FIGURE 22. Aulacus froggatti sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus.

FIGURE 23. Aulacus fuscicornis Cameron, holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) dorsal mesosoma; (F) lateral mesosoma; (G) dorsal hind coxae. Aulacus fuscicornis Cameron, male, (H) ventral male genitalia.

FIGURE 24. Aulacus glorious sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) frontal mandibles; (E) dorsal mesoscutum; (F) lateral mesosoma.

FIGURE 25. Aulacus hackeri sp. nov., holotype female, (A) lateral habitus; (B) dorsal habitus; (C) dorsal hind coxae.

FIGURE 26. Aulacus insularis sp. nov., paratype male, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head and mesosoma (part); (E) dorsal mesosoma; (F) lateral mesosoma.

FIGURE 27. Aulacus jamberoo sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) fronto-dorsal head; (D) frontal head; (E) dorsal mesosoma; (F) lateral mesosoma; (G) dorsal metasomal T1; (H) dorsal hind coxae.

FIGURE 28. Aulacus kittelae sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head and mesosoma.

FIGURE 29. Aulacus kiwarrakensis sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) lateral head; (D) dorsal mesosoma; (E) lateral mesosoma; (F) dorsal hind coxae.

FIGURE 30. Aulacus leai sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) lateral head; (D) dorsal mesosoma; (E) lateral mesoscutum; (F) dorsal hind coxae.

FIGURE 31. Aulacus longiventris (Kieffer), female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) dorsal mesosoma; (F) lateral mesosoma; (G) dorsal hind coxae. Aulacus longiventris (Kieffer), male, (H) ventral male genitalia.

FIGURE 32. Aulacus mareebaensis sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head, lateral propodeum and frontal mesosoma; (D) lateral head and dorsal mesoscutum; (E) lateral mesosoma; (F) dorsal hind coxae.

FIGURE 33. Aulacus minutus Crosskey, female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) dorso-lateral head and propodeum; (E) dorsal mesoscutum; (F) lateral pronotum and mesoscutum; (G) outer lateral hind coxa; (H) dorsal hind coxae.

FIGURE 35. Aulacus nebo sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorso-lateral head, pronotum and mesoscutum; (D) dorsal hind coxae.

FIGURE 36. Aulacus neboissi sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) dorsal mesosoma; (F) lateral pronotum and mesoscutum.

FIGURE 37. Aulacus obcordellus sp. nov., paratype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) dorsal mesosoma; (F) dorso-lateral pronotum and mesoscutum.

FIGURE 38. Aulacus pallidicaudis (Cameron), holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) dorsal mesosoma; (F) lateral mesosoma; (G) frontal head; (H) dorsal metasomal T1.

FIGURE 41. Aulacus quickei sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) lateral head, pronotum and mesoscutum; (D) dorsal head; (E) dorsal mesosoma; (F) dorsal metasomal T1 and T2.

FIGURE 42. Aulacus rieki sp. nov., paratype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) dorso-lateral head.

FIGURE 43. Aulacus scitulus sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head, pronotum and mesoscutum; (D) lateral head, pronotum and mesoscutum; (E) dorsal hind coxae; (F) frontal head.

FIGURE 45. Aulacus smithi sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) lateral head; (D) dorsal head; (E) lateral mesosoma; (F) dorsal hind coxae.

FIGURE 46. Aulacus tasmanicus sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) dorsal mesosoma; (E) lateral mesosoma; (F) dorsal propodeum and hind coxae.

FIGURE 47. Aulacus tiernyi sp. nov., paratype female, except A holotype (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) dorsal mesoscutum; (E) lateral mesosoma; (F) dorsal hind coxae.

FIGURE 48. Aulacus truncatus (Kieffer), holotype female, (A) lateral habitus; (B) lateral metasoma; (C) dorsal habitus (Images ©The Trustees of the Natural History Museum, London)).

FIGURE 50. Aulacus vespiformis (Kieffer), holotype male, (A) dorsal habitus; (B) lateral habitus (Images ©The Trustees of the Natural History Museum, London).

FIGURE 51. Aulacus walkeri sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) dorsal mesoscutum; (E) lateral mesosoma; (F) dorsal hind coxae.

FIGURE 53. Aulacus willamsi sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) lateral head; (E) dorsal mesoscutum; (F) dorsal mesosoma; (G) dorsal hind coxae; (H) inner lateral right hind coxa (paratype).

FIGURE 54. Aulacus wrightae sp. nov., holotype female, (A) dorsal habitus; (B) lateral habitus; (C) dorsal head; (D) frontal head; (E) dorsal mesosoma; (F) dorsal hind coxae.