Verbascum blattaria L., Sp. Pl. 1: 178. 1753.

Khamar, Hamid, Oualidi, Jalal El, Touhami, Amina Ouazzani & Civeyrel, Laure, 2023, Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa, PhytoKeys 225, pp. 115-152 : 115

Verbascum blattaria L., Sp. Pl. 1: 178. 1753.

Blattaria alba = Blattaria alba Mill., Fig. Pl. Gard. Dict. 1: 45. 1760.

Verbascum glabrum = Verbascum glabrum Mill., Gard. Dict. ed. VIII: n. 8 1768.

Verbascum cordatum = Verbascum cordatum Desf., in Fl. Atl. 1: 186. 1798.

Verbascum repandum = Verbascum repandum Willd., Enumeratio Plantarum: 226. 1809.

Thapsus blattaria ≡ Thapsus blattaria (L.) Raf., in Fl. Tellur. 4: 89. 1838.

Verbascum blattaria albiflorum = Verbascum blattaria var. albiflorum G.Don, in Gen. Hist. 4: 497 1838. ≡ Verbascum blattaria f. albiflora (G.Don) House, in Bull. New York State Mus. Nat. Hist. 243-244: 45. 1923.

Verbascum blattaria albiflorum = Verbascum blattaria var. albiflorum Kuntze, in Revis. Gen. Pl. 2: 468.1891, nom. illeg.

Blattaria vulgaris = Blattaria vulgaris Fourr., in Ann. Soc. Linn. Lyon, N. S. 17: 125. 1869.

Verbascum blattaria brevipedicellatum = Verbascum blattaria L. var. brevipedicellatum Halácsy. in Ost. bot. Zeitschr. XLII: 419. 1892. Type: [Greece]. Insula Thasos. Limenas, in pratis arenosis in oliveto. 04 June 1891, P. Sintenis & J. F. N Bornmüller 655 (Lectotype, designated here: LD [LD1393506 image!]; isolectotypes JE [JE00012328 image!], K [K000806399 image!], B [B100278360 image!, B100278359 image!]. [image of lectotype available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.ld1393506]. (2)

Verbascum blattaria crenatum = Verbascum blattaria var. crenatum Rouy in Rouy & Foucaud, Fl. France 11: 10. 1909.

Verbascum rhinanthifolium = Verbascum rhinanthifolium Davidov in Trav. Soc. Bulg. Sci. Nat. 8: 101. 1915.

Verbascum carduifolium = Verbascum carduifolium Murb. ex. Hayek, Repert. Spec. Nov. Regni Veg. Beih. 30(2): 131. 1929. ≡ Verbascum blattaria var. carduifolium Murb., in Lunds Univ. Arsskrift, 2n.f., 29 (2): 567. 1933. Type: [Greece]. Malakasi; in vinetis, 17 June 1896, P. Sintenis, 632. (leolotype, designated here: B [B100278358 image!]; isolectotypes: WU [WU0126534 image!], LD [LD1364911 image!, LD1395023 image!, LD1392908 image!, LD1394623 image!]). [image of lectotype available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.b_10_0278358]. (3)

Verbascum blattaria gracilipes = Verbascum blattaria var. gracilipes Murb. in Lunds Univ. Arsskrift, 2n.f., 29 (2): 566. 1933. Type: [Greece]. Volo: Portaria, in vinetis, 02 September 1896, P. Sintenis 1301 (Lectotype, designated here: B [B100278357 image!]; isolectotypes: JE [JE00012330 image!, JE00012331 image!]; WU [WU0126494 image!]). [Greece]. In valle Tempe: prope Papapuli 28-31 Jully 1913, B. Tuntas, 1954 (Residual syntype: WU [WU0126493]!; In valle Tempe: prope Papapuli 28-31 July 1913, B. Tuntas, 1955 (Residual syntype: WU [WU0126492 image!]). [image of lectotype available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.b_10_0278357]. (4)

Verbascum blattaria brachycalyx = Verbascum blattaria var. brachycalyx Murb. in Lunds Univ. Arsskrift, 2n.f., 29 (2): 567. 1933. Type: [Greece]. Insula Zakynthos (Zante): in humidis montis Skopos; reg. infer, 4 May 1926, J. F. N. Bornmüller 1177 (Lectotype, designated here: B [B100278364 image!]; isolectotypes: B [B100278363 image!]). [Greece]. Insula Kephalonia, in olivetis ad Argostoli, 23 May 1926, J. F. N. Bornmüller 1174 (Residual syntype: JE [JE00012329 image!], S [S10-26942 image!, S12-12718 image!]). [Greece]. Insula Zakynthos (Zante), in herbidum montis Skopos, 04 May 1926, J. F. N. Bornmüller 1177 (Residual syntype: S [S10-26943 image!]). [GREECE]. Insula Zakynthos (Zante); in olivetis 23 May 1926, J. F. N. Bornmüller 1174 (Residual syntype: B [B100278367 image!]). [Greece]. Insula Zakynthos (Zante); in olivetis, 02 May 1926, J. F. N. Bornmüller 1175 (Residual syntype: B [B100278366 image!]). [image of lectotype available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.b_10_0278364]. (5)

Type.

(lectotype was designated by Huber-Morath 1971, pg. 143): LINN [Herb. Linn. No. 242.6.]).

Notes.

Original material is kept in the Linnaean Herbarium at the Linnean Society of London; an image of the lectotype is available at http://linnean-online.org/1836/.

(2) No specimens were cited in the protologue ( Halácsy 1892) of Verbascum blattaria L. var. brevipedicellatum . However, according to Murbeck (1933) the description of this taxon was based on material collected by Sintenis and Bornmüller from 1883 to1892 in Greece. We have located five sheets that have been lodged at the following four Herbs: LD (LD1393506), JE (JE00012328), K (K000806399), and B (B100278360, B100278359), the labels’ data of which correspond to those reported by Murbeck (1933). Hence, these specimens are eligible and can be considered as original material. The sheet LD1393506 preserved at Herb. LD is here selected as the lectotype of the name Verbascum blattaria L. var. brevipedicellatum .

(3) The name of Verbascum carduifolium was first published by Murbeck (1925: 168), but without a description. Four years later, Hayek (1929) gave the description of V. carduifolium Murb. ex. Hayek and mentioned a locality “The.” [as an abbreviation of Thessalía]. Four years later, the taxonomic status of this species has been assessed by Murbeck (1933: 567) and he regarded it as a variety under Verbascum blattaria L. In addition to the morphological description, Murbeck (1933) cited an element collected by P. Sintenis viz. "In monte pindo Malakasi, in pratis supra pagum. determ. Leg. P. Sintenis 17 /6/ 1896, n. 632" and he indicated the herbaria (B, WU, LD) that housed the specimens he surveyed. We located six sheets, all of them from the locality given in the protologue, belonging to the Sintenis collections: B (Barcode B100278358), WU (WU0126534) and LD (LD1364911, LD1395023, LD1392908, LD1394623) at Herb. The sheet B100278358 conserved at Herb. B is here designated as lectotype for the name V. carduifolium .

(4) In the protologue of Verbascum blattaria var. gracilipes , Murbeck (1933) cited 11 gatherings, but he does not provide the name of the herbarium where the original material has been deposited. Six sheets from three different Herbs (B, JE, WU) were traced only: B (Barcode B100278357), JE (JE00012330, JE00012331) and WU (WU0126492, WU0126493, WU0126494). All of these specimens are consistent with the location and diagnosis given in the protologue and can be considered as original material. Since no specimen-type has been designated, all of these specimens can be considered syntypes under Art. 9.6 of the ICN ( Turland et al. 2018). The specimen B100278357 preserved at Herb. P is selected here as the lectotype for the name of this variety.

(5) In describing Verbascum blattaria var. brachycalyx , Murbeck (1933) referred to eight collections made in five different regions in Greece, but he did not designate a single type for the name. We traced eight specimens in Herbs. B (B100278364, B100278363, B100278367, B100278366), S (S10-26942, S10-26943, S12-12718) and JE (JE00012329) which can be considered original material. We here designate the specimen B100278364 preserved at Herb. B as the lectotype of the name V. blattaria var. brachycalyx since it is well preserved.