Republica weatbrooki Archibald & Cannings, 2021

Archibald, S. Bruce & Cannings, Robert A., 2021, A new genus and species of Euphaeidae (Odonata, Zygoptera) from the early Eocene Okanagan Highlands locality at Republic, Washington, U. S. A., Zootaxa 4966 (3), pp. 392-400 : 396-397

Republica weatbrooki Archibald & Cannings , new species

Figure 1

Diagnosis. As for the genus.

Material. Holotype: SR 06-59 - 08 , collected by Alex Weatbrook, 24.v.2006. An almost complete forewing (presumed, see above) with small parts of the posterior margin missing in the apical half, some regions indistinctly preserved between main veins. Housed in the Stonerose Interpretive Center collections.

Description. Holotype, forewing (see above) SR 06-59-08. 40.7 mm long, 10.4 mm maximum width at about mid-wing; nodus 14.0 mm from base, pterostigma 33.3 mm from base; lightly infuscate as preserved in the proximal three quarters and portions of apical quarter (possibly an artefact of preservation); without petiole. At least 28 antenodal crossveins, at least 26 Ax2 to nodus (C–ScP space), surely many more, space not completely preserved; some pairs originating on ScP join; ScP–RA space: 19 preserved (minimum, space incompletely preserved), some not aligned with those in C–ScP space; Ax1, Ax2 distinctly bolder than accessory antenodal crossveins; one crossvein between Ax1, Ax 2 in costal space. Postnodal crossveins numerous, dense, some branching, joining, some joined by crossveins, C–RA, RA–RP1 crossveins preserved not aligned; C–RA space distal to pterostigma maximum three cells wide. Wing very dense with crossveins throughout (where preserved: some distal regions obscured by preservation). Region where oblique brace vein might be, if present, is not preserved. Pterostigma: 4.0 mm long, about 0.7 mm wide; basal, distal sides oblique. Main veins apical to MA mostly rather straight, only gently curve to hind margin near terminus; MA undulate, smoothly curving twice toward anterior margin; CuA somewhat more deeply curved, ends on margin about mid-wing. Identities of RP1, IR2, at apex not clear by preservation; about 18 secondary longitudinal veins reach margin as preserved; three between MA, MP, probably more; none detected between MP, CuA; between CuA, posterior margin. Quadrangle+sub-quadrangle length about twice height (MP+CuA dividing them not preserved); thick discoidal bracket (“db”, Fig. 1 ) with reverse obliquity; subquadrangle with at least two crossveins (none preserved in quadrangle). Posterior base of wing (mostly posterior to subquadrangle, region basal to this) narrow, few cells about twice high as long.

Etymology. The specific epithet is a patronymic formed from the surname of Alex Weatbrook, who found and donated the holotype, recognising his contribution.

Range and age. Tom Thumb Tuff Member of the Klondike Mountain Formation, exposure B4131 at Republic, Washington, USA; latest Ypresian .

Figures

FIGURE 1. Republica weatbrooki Holotype SR 06-59-08. A, photograph; B, drawing. Both to scale, 5 mm; db, discoidal bracket, see text.