Heliobolus Fitzinger, 1843

Marques, Mariana P., Ceríaco, Luis M. P., Blackburn, David C. & Bauer, Aaron M., 2018, Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola Atlas of Historical and Bibliographic Records (1840 - 2017), Proceedings of the California Academy of Sciences 65, pp. 1-501 : 213-214

Genus Heliobolus Fitzinger, 1843

Heliobolus lugubris ( Smith, 1838) BUSHVELD LIZARD

Lacerta lugubris Smith 1838:93 . Syntypes: BMNH 1946.8.6.37-45 (collector A. Smith). Type locality: “District immediately beyond the northern frontier of the [Cape] Colony,” South Africa. Eremias lugubris : Bocage (1867b:221, 1895a:31), Boulenger (1905:111; 1921:239), Monard (1937b:75),

Mertens (1938a:437). Lampreremias lugubris : Szczerbak (1975:33). Heliobolus lugubris : Branch (1998:161), Bates et al. (2014:160), Ceríaco et al. (2016a:37, 56), Conradie et al.

(2016:24).

Global conservation status (IUCN): Not Evaluated.

Global distribution: Widely distributed, from southern Angola over the Kalahari sands through Namibia, Botswana, southwestern Zimbabwe, southern Mozambique and South Africa.

Ocurrences in Angola (Map 166): The species occurs broadly in southern Angola.

Benguela: “ Sighting in Lobito ” [-12.35000,

13.55000] ( Monard 1937b:75); “Catumbella”

[-12.43333, 13.55000] ( Bocage 1895a:31);

“Quissange” [-12.43333, 14.05000] ( Bocage

1895a:31); “Benguella” [-12.58333, 13.41667]

( Bocage 1867b:221, 1895a:31; Szczerbak

1975:33); “Dombe” [-12.95000, 13.10000]

( Bocage 1895a:31); “Cubal” [-13.03333,

14.25000] ( Mertens 1938a:437); “Huxe,

Benguella” [-12.71667, 13.20000] (Boulenger

1921:243). Namibe: “Maconjo” [-15.01667,

13.20000] ( Bocage 1895a:31; Ceríaco et al.

2016b:56); “Namibe-Lubango road, 2 km E of

Mangueiras, south side of the road”

[-15.04361, 13.16000] (Ceríaco et al.

MAP 166. Distribution of Heliobolus lugubris in 2016a:37); “Capangombe” [-15.10000, Angola.

13.15000] ( Bocage 1895a:31; Ceríaco et al. 2016a:56); “Konondoto, Mossamedes” [-15.20000, 12.15000] (Boulenger 1921:243; Ceríaco et al. 2016a:56); “Namibe-Lubango road, 2 km E of Mangueiras, south side of the road” [-15.03333, 13.16000] (Ceríaco et al. 2016a:37); “Mossamedes” [-15.78333, 12.06667] ( Szczerbak 1975:33). Cunene: “Ponang Kuma” [-17.05000, 14.65000] (Boulenger 1921:243); “Mupanda” [-17.13333, 15.76667] ( Monard 1937b:75). Cuando Cubango: “Cuito basin (63)” [-17.93611, 21.10269] ( Conradie et al. 2016:9, 12, 24). Undetermined Locality: “Cuanza R.” ( Boulenger 1905:111).

Taxonomic and distributional notes: This widespread species has never been investigated in detail and it is likely that there is regional differentiation and possibly even cryptic taxa within H. lugubris as currently construed.