Lubuya Horton, 1972

Marques, Mariana P., Ceríaco, Luis M. P., Blackburn, David C. & Bauer, Aaron M., 2018, Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola Atlas of Historical and Bibliographic Records (1840 - 2017), Proceedings of the California Academy of Sciences 65, pp. 1-501 : 244-245

Genus Lubuya Horton, 1972

Lubuya ivensii (Bocage, 1879) IVENS’ SKINK Euprepes Ivensi Bocage 1879a:97 . Syntypes: MBL (3 specimens) numbers not known (collectors H. Capello and R. Ivens), destroyed by fire 18 March 1978. Type locality: “Bihé, dans l’intérieur de Benguella” [= Bié], Bié Province, Angola.

Mabuya ivensi septemlineata ( Laurent 1964a:77, fig. 23). Holotype: MD 5427 (collector A. Barros Machado). Type locality: “Alto Chicapa, Lunda” [Lunda Sul Province, Angola].

Euprepes Ivensi : Bocage (1879b:95).

Mabuia ivensii : Boulenger (1887:197).

Lygosoma Ivensii : Bocage (1895a:48, 1897a:196).

Mabuya ivensii : Monard (1937b:86), Manaças (1963:233), Branch and Haagner (1993).

Lubuya ivensii : Horton (1972:17), Metallinou et al. (2016:4).

Trachylepis ivensii : Wagner et al. (2012:39).

Global conservation status (IUCN): Not Evaluated.

Global distribution: The species is widely distributed through the drainage systems of Angola, extending into the adjacent regions of the southern Democratic Republic of Congo and Zambia. Ocurrences in Angola (Map 204): The species is known from central-east Angola.

Lunda Sul: “Alto Cuílo” [-10.01667,

19.55000] ( Laurent 1964a:77; Branch and

Haagner 1993:112; Wagner et al. 2012:37);

“Alto Chicapa” [-10.88333, 19.23333] ( Laurent 1964a:77; Branch and Haagner 1993:112;

Wagner et al. 2012:37); “Dala” [-11.03333,

20.20000] ( Monard 1937b:86; Laurent

1964a:78; Branch and Haagner 1993:112;

Wagner et al. 2012:37). Malanje: “ Rio Loando” (R. Luando, près de Mongoa; Luando

River, tributary of the Cuanza River, near Mongoa )” [-11.55000, 18.15000] ( Bocage

1879b:95; Laurent 1964a:78; Branch and

Haagner 1993:112; Wagner et al. 2012:37);

“entre Malanje et Pungo Andongo” ( Laurent

1964a:78). Moxico: “source of the Calombe MAP 204. Distribution of Lubuya ivensii in Angola. River (Luso)” [-11.83333, 19.93333] ( Manaças 1963:233; Horton 1972:17); “Luso (= Luena)” [-11.78333, 19.91667] ( Horton 1972:17; Branch and Haagner 1993:112; Wagner et al. 2012:37). Bié: “Bihé, dans l’intérieur de Benguella” [-12.38333, 16.95000] ( Bocage 1879a:97); “Benguella” (“Bihé” according to the original publication fide Bocage 1879a:97) [-12.38333, 16.95000] ( Boulenger 1887:197; Horton 1972:17). Huambo: “ Huambo ” [-12.76667, 15.73333] ( Horton 1972:17; Branch and Haagner 1993:109, 112; Wagner et al. 2012:37); “R. Cuando River, affluent du Cunene ” [-13.38333, 15.71667] ( Laurent 1964a:78; Branch and Haagner 1993:112; Wagner et al. 2012:37). Undetermined locality: “Quando” ( Bocage 1895a:48, 1897a:196; Branch and Haagner 1993:112); “sur les bords du Quanza et du Quando (margens do Quanza e do Quando)” ( Bocage 1895a:49, 1897a:196); “R. Cuanza” ( Laurent 1964a:78; Branch and Haagner 1993:112; Wagner et al. 2012:37).

Taxonomic and distributional notes: Boulenger (1887) incorrectly restricted Bocage’s type locality to “ Benguela,” an action later accepted by Horton (1972), Branch and Haagner (1993), and Wagner et al. (2012). The species was described in Euprepes but later moved by Bocage (1895a) to Lygosoma . Horton (1972) described the monotypic genus Lubuya to accommodate M. ivensii , differentiating it from Mabuya as then construed. Greer (1977) suggested that Lubuya should return to Mabuya , or by implication to Trachylepis , the name subsequently allocated to African mabuyine skinks (Bauer 2003). However, Metallinou et al. (2016) ressurected Lubuya to accommodate T. ivensii , which they found to be the distantly related sister to Eumecia . Monard (1937b) was the first to recognize that specimens from northeastern Angola had an extra white line on the lower flank and Laurent (1964a) subsequently erected Mabuya ivensi septemlineata to recognize this form; its validity has not been critically assessed.