Polydrepanini, Jeekel, 1968

Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha & Sudhikumar, Ambalaparambil Vasu, 2021, Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae), Zootaxa 5068 (4), pp. 485-516 : 509

Key to the accepted genera of Polydrepanini , based on gonopodal characters

1(2) Femorite clearly twisted ( Figs 9–12 , 16–20 , 45–53 ).......................................................... 3

2(1) Femorite untwisted ( Figs 3–8 , 28–44 , 55–57 , 60–64 , 67, 68 , 84–87 )............................................. 5

3(4) Solenophore particularly simple, two upright lobes either side of a higher and suberect solenomere, the latter devoid of a basal curve/loop ( Figs 45, 46 )....................................................................... Dasypharkis

4(3) Solenophore much more complex, strongly coiled; solenomere with a basal curve/loop ( Figs 9–12 , 17–20 , 47–53 )................................................................................................. Polydrepanum

5(6) Solenomere with neither a distinct basal loop nor a protecting lobe at its base...................................... 7

6(5) Solenomere with both a distinct basal loop and a protecting lobe at its base....................................... 9

7(8) Solenophore relatively small, but clearly twisted ( Figs 67, 68 )....................................... Xiphidiogonus

8(7) Solenophore usually large, varied ( Figs 28–44 , 84–87 )............................................... Delarthrum

9(10) Solenophore cup-shaped, enlarged distad ( Figs 55–57 )................................................ Pocockina

10(9) Solenophore otherwise................................................................................ 11

11(12) Both solenophore and solenomere either suberect ( Figs 3–5 ) or subcircular ( Figs 6–8 ).................. Grammorhabdus

12(11) Both solenophore and solenomere strongly coiled, the former with two distomesal processes (a and b), b being unusually large and shield-like ( Figs 60–64 )................................................................... Telodrepanum

Figures

FIGURES 9–12. Polydrepanum horridum Golovatch, 1984. Right gonopod, lateral, dorsal, mesal and ventral views, respectively.After Golovatch (1984). Abbreviations: a, apicolateral spine; k, basal loop of solenomere; lo, mesal lobe; sl, solenomere; sph, solenophore. Scale bar: 0.4 mm.

FIGURES 16–20. Polydrepanum tamilum Carl, 1932. 16. Male leg 2, lateral view. 17. Entire left gonopod, mesal view. 18. Distal half of left gonopod, lateral view. 19. Entire right gonopod, mesal view. 20. Distal half of right gonopod, sublateral view. 16–18: after Carl (1932); 19, 20: after Jeekel (1980b). Abbreviations: a and b, dorsal distofemoral processes; ad, adenostyle; lo, lateral protecting lobe/shield; k, basal loop of solenomere; sl, solenomere; sph, solenophore. Drawn not to scale.

FIGURES 45, 46. Dasypharkis rugulosa (Carl, 1932) (= Orthomorpha rugulosa Carl, 1932). Entire left (45) and right (46) gonopod, mesal and lateral views, respectively. After Jeekel (1980b). Abbreviations: a, lateral distofemoral spine; sl, solenomere; sph, solenophore. Drawn not to scale.

FIGURES 47–51. Polydrepanum lamprum (Chamberlin, 1920) (= Orthomorpha lampra Chamberlin, 1920). 47. Entire right gonopod, mesal view. 48, 49. Distal half of right gonopod, sublateral and mesal views, respectively. 50. Entire left gonopod, mesal view. 51. Distal half of left gonopod, ventral view. 47–49: after Jeekel (1972);: 50, 51: after Carl (1932). Abbreviations: a, mesal distofemoral spine; b, lateral distofemoral spine; c, ventromesal distofemoral process; k, basal loop of solenomere; lo, lateral protecting lobe; sl, solenomere; sph, solenophore. Drawn not to scale.

FIGURES 52, 53. Polydrepanum granuliferum (Attems, 1936) (= Sundanina granulifera Attems, 1936). Entire right gonopod, lateral and mesal views, respectively.After Golovatch (1984). Abbreviations: a, mesal distofemoral spine; b, lateral distofemoral spine; k, basal loop of solenomere; lo, lateral protecting lobe; sl, solenomere; sph, solenophore. Scale bar: 0.5 mm.

FIGURES 1–5. Grammorhabdus asperrimus Carl, 1932. 1. Male leg 1, lateral view. 2. Male leg 2, lateral view. 3. Entire left gonopod, mesal view. 4, 5. Distal half of left gonopod, lateral view. 1–4: after Carl (1932), 5: after Jeekel (1980B). Abbreviations: a and b, apicomesal spines; ad, adenostyle; ca, cannula; cx, coxite; fe, femorite; k, basal loop of solenomere; lo, lateral lobe; pfe, prefemoral portion; sl, solenomere; sph, solenophore. Drawn not to scale.

FIGURES 6–8. Grammorhabdus fissus (Sankaran & Sebastian, 2018). 6, 7. Right gonopod, ventral and mesal views, respectively. 8. Distal half of right gonopod, lateral view. After Sankaran and Sebastian (2018). Abbreviations: a and b, apicomesal lobes; ad, adenostyle; ca, cannula; cx, coxite; fe, femorite; k, basal loop of solenomere; lo, lateral lobe; pfe, prefemoral portion; sl, solenomere; sph, solenophore; su, distofemoral sulcus. Scale bars: 0.5 mm (6, 7) and 0.2 mm (8).

FIGURES 27–30. Delarthrum obscurum Attems, 1936. 27. Male femur 1, lateral view. 28. Entire right gonopod, lateral view. 29, 30. Distal half of right gonopod, lateral and mesal views, respectively. After Attems (1936). Abbreviations: a, mesal distofemoral process; ad, adenostyle; sl, solenomere; sph, solenophore. Drawn not to scale.

FIGURES 31–33. Delarthrum spiniger (Attems, 1936) (= Armolites spiniger (Attems, 1936)). 31. Entire left gonopod, mesal view. 32, 33. Distal half of left gonopod, ventral and dorsal views, respectively. After Golovatch (1984). Abbreviations: a, mesal distofemoral process; sl, solenomere; sph, solenophore. Scale bar: 1.0 mm.

FIGURES 34–36. Delarthrum unicolor (Attems, 1936) (= Martensosoma unicolor (Attems, 1936)). 34. Entire right gonopod, mesal view. 35, 36. Distal half of right gonopod, lateral and submesal views, respectively.After Attems (1936). Abbreviations: sl, solenomere; sph, solenophore. Drawn not to scale.

FIGURES 37–39. Delarthrum kuznetsovi (Golovatch, 1994) (= Nepalomorpha kuznetsovi Golovatch, 1994). 37. Entire left gonopod, mesal view. 38, 39. Distal half of left gonopod, dorsal and lateral views, respectively. After Golovatch (1994a). Abbreviations: sl, solenomere; sph, solenophore. Scale bar: 0.5 mm.

FIGURES 40, 41. Delarthrum hingstoni (Carl, 1935) (= Orophosoma hingstoni (Carl, 1935)). 40. Entire right gonopod, mesal view. 41. Distal half of left gonopod, lateral view. After Carl (1935). Abbreviations: a, dorsal distofemoral process; p, basal prefemoral process; sl, solenomere; sph, solenophore. Drawn not to scale.

FIGURES 42–44. Delarthrum longisetum (Golovatch, 1994) (= Parorthomorpha longiseta Golovatch, 1994). 42. Entire left gonopod, mesal view. 43, 44. Distal half of left gonopod, dorsal and lateral views, respectively. After Golovatch (1994a). Abbreviations: sl, solenomere; sph, solenophore. Scale bar: 0.2 mm.

FIGURES 54–57. Pocockina pilifera (Pocock, 1895). 54. Male leg 1, lateral view. 55. Entire right gonopod, mesal view. 56, 57. Distal half of left gonopod, lateral and mesal views, respectively. After Jeekel (1965). Abbreviations: ad, adenostyle; fe, femorite; k, basal loop of solenomere; lo, mesal protecting lobe; sl, solenomere; sph, solenophore. Drawn not to scale.

FIGURES 58–60. Telodrepanum badaga Carl, 1932. 58. Male leg 1, lateral view. 59. Male leg 2, lateral view. 60. Distal half of left gonopod, mesal view. After Carl (1932). Abbreviations: a and b, mesal distofemoral processes; ad, adenostyle; lo, lateral protecting lobe; sl, solenomere; sph, solenophore. Drawn not to scale.

FIGURES 61–64. Telodrepanum sp. from Dharwar, India. Entire right gonopod, dorsomesal, mesal, subventral and sublateral views, respectively. After Golovatch (1984). Abbreviations: a and b, mesal distofemoral processes; k, basal loop of solenomere; lo, lateral protecting lobe; p, coxital process; sl, solenomere; sph, solenophore. Scale bar: 1.0 mm.

FIGURES 65–68. Xiphidiogonus spinipleurus Carl, 1932. 65. Male leg 1, lateral view. 66. Male leg 2, lateral view. 67. Entire right gonopod, mesal view. 68. Distal half of right gonopod, lateral view. After Carl (1932). Abbreviations: a, b, c and d, distofemoral processes; ad, adenostyle; fe, femorite; k, basal loop of solenomere; sl, solenomere; sph, solenophore. Drawn not to scale.

FIGURES 81–87. Delarthrum anomalans sp. nov., male paratypes. 81. Legs 1, lateral view. 82. Legs 2, lateral view. 83. Leg 26 (anterior pair on body ring 16), lateral view. 84–87. Entire right gonopod, mesal, submesal, mesal and subventral views, respectively. Abbreviations: a and b, distofemoral processes; fe, femorite; sl, solenomere; sph, solenophore; z, dorsobasal peg.