Anabarhynchus tribulationensis Ferguson

Ferguson, David J., Irwin, Michael E. & Yeates, David K., 2013, New species of Anabarhynchus Macquart (Diptera: Therevidae) from arid and monsoon tropical Australia, Zootaxa 3680 (1), pp. 55-95 : 88-91

Anabarhynchus tribulationensis Ferguson sp. n.

( Figures 26 , 27 , 28 )

Type material. Holotype: Male. AUSTRALIA: Queensland: Cape Tribulation. N. Qld., 8.IV.1982, D.E. Hardy ( ANIC _29:028797) ( ANIC). Condition: Flagellum missing, pined with micro pin dorsally to pith block, end of abdomen removed for dissection after photography, reasonable condition.

Diagnosis. Frons raised; male frons width at anterior ocellus 2.6x ocellus; ocellar; mid frons with dark-brown marks angled towards upper frons near meeting at mid-line; lower frons setae arrange above and as wide as antenna bases. Scutum grey tones with indistinct lines. Wing cell m3 open. Postspiracular pile present; katepisternum and prosternal furrow without pile. Forefemur, 2 pd, 1–2 pv; mid-femur 1 pd, 4–7 pv, 2 av; hind femur 1–2 av; femora basally dark apically yellow-brown; all femora with appressed pale pile dorsally, ventrally semi-erect, admix with sparse short black setae predominantly to apical half. Male abdominal integument yellow-brown.

Description. Male: Body length: 7.5mm. Wing length: 6mm. Head. Integument black. Frons slightly raised; frons width at anterior ocellus 2.6x ocellus; ocellar tubercle flat; antennal base positioned low on frons, face and lower frons slightly protruding; parafacials and lower frons bright grey, middle lower frons brown-grey; mid-frons with blackish brown upside-down ‘V’ mark; frons setae in two rows; lower frons setae arrange above and as wide as antenna bases; lower frons setae thirds the length of scape. Scape length 2x width; scape and pedicel grey, flagellum missing. Occiput convex generally grey pubescence, margin of eye yellowish- brown, several indistinct rows of black macrosetae 29 each side; postocciput to gena grey with long, pale, hair-like pile. Palpus pale yellow sparsely supplies with pale hair like pile; labellum brown grey; prementum without dark setae. Thorax. Integument black. Scutal chaetotaxy black (pairs): np, 4; sa, 2; pa, 1; dc, 2; sc, 2. Scutum with thin brown dorsal line within broader grey band, margined with indistinct brownish-grey line, beside broader pale grey line, irregular brownishgrey marks margin laterally raised grey areas pre and post-transverse suture; scutal surface with sparse short black setae. Postspiracular pile present; katepisternum and prosternal furrow without pile; pleuron and coxae with grey pubescence; coxae with elongate pale pile admixed with black macrosetae; posterior coxa 3 with pile to anterior margin. Wing. Cell m3 open; hyaline with brown tint; brown veins; stigma brown infuscate along margin. Costal setae beyond humeral cross-vein biserially arranged. Haltere. Pedicel buff-white; scabellum buff-white. Legs.

Forefemur with 2 pd, 1–2 pv weak; mid-femur 1 pd, 4–7 pv weak, 2 av; hind femur 1–2 av macrosetae; forefemur basal two-thirds dark brown apically third pale brown; mid-femur pale brown; hind femora basally extending along dorsal dark brown, apical and ventrally yellow-brown; all femora with appressed pale pile dorsally, ventrally semierect, admix with sparse short black setae predominantly to apical half; all tibia pale brown darkening apically. Abdomen. Integument yellow-brown; slightly laterally compressed; anterior bands broadly yellow-brown, anteriorly viewed matte yellow-brown, with weak appressed dark pile; tergites 2–7 posterolaterally bright grey with appressed pale pile, tergites 2–4 posterior marginal bands when viewed posteriorly bright white, anteriorly grey; tergites 5–7 with black setae; epandrium yellow-brown; gonocoxite yellow-brown. Terminalia. Epandrium ( Fig. 28 a), almost three times wide as long, slightly narrowing posteriorly, setae on posterior lateral edges. Gonocoxite ( Fig. 28 b), semi-spherical slightly longer than wide when viewed ventrally; posterior ventral edge with broad slightly ventrally directed flange. Joined along hypandrium. Gonocoxal apodeme extending just beyond lateral margin, moderately sclerotised. Inner gonocoxal process longer than gonostylus, inwardly curved and ventrally directed several robust setae on sub-apical inner ventral edge. Gonostylus dorsally directed with long thin setae evenly distributed over dorsal surface and inner middle ventral surface directed inward, apical end reflexed dorsally. Ventral lobe absent. Aedeagus ( Fig. 28 c, d): distiphallus strongly curved ventrally. Parameral sheath sclerotised. Dorsal apodeme of parameral sheath broadly triangular; apical ends dorsally directed. Ventral apodeme basally narrow, broadly widening apically. Lateral ejaculatory apodeme dorsally broad, triangular shape. Ejaculatory apodeme, cylindrical, sub-apical flanges laterally.

Female. Unknown.

Etymology. The specific epithet is derived from ‘Cape Tribulation’ the geographic location where the type specimens were collected.

Comments. Know from a single male collected in April from Cape Tribulation, North Queensland.

Keys to couplet 88 in Lyneborg (2001) and readily separated from A. boharti Lyneborg by being a small species, the abdominal integument yellow brown with anterior bands yellow brown and distinctive dark up-sidedown ‘V’ to frons.

Can be separated from A. ravenshoensis sp. n., by the frons being slightly raised, lacking rouges striation to upper frons. Mid-frons with blackish brown upside-down ‘V’ mark. Lower frons setae being arrange above and as wide as antenna bases. Forefemur lack av macrosetae and mid-femur have several pv macrosetae.

Anabarhynchus tribulationensis sp. n., is closely related to both A. ewamin sp. n., and A. ravenshoensis sp. n.; subsequently placed with the kroeberi species-group. For femoral macrosetae differences within the kroeberi species-group see Table 1.

Figures

FIGURE 26. Anabarhynchus tribulationensis sp. n., holotype male, (ANIC _ 29: 028797): lateral view.

FIGURE 27. Anabarhynchus tribulationensis sp. n., holotype male, (ANIC _ 29: 028797): oblique view.

FIGURE 28. Anabarhynchus tribulationensis sp. n., holotype male terminalia (ANIC _ 29: 028797): A, epandrium, left dorsal view, right ventral view; B, atrium left dorsal view, right ventral view; C, aedeagus, dorsal view; D, aedeagus, lateral view; Scale line = 0.5 mm.