Anabarhynchus ewamin Ferguson

Ferguson, David J., Irwin, Michael E. & Yeates, David K., 2013, New species of Anabarhynchus Macquart (Diptera: Therevidae) from arid and monsoon tropical Australia, Zootaxa 3680 (1), pp. 55-95 : 61-65

Anabarhynchus ewamin Ferguson sp. n.

( Figures 5 , 6 , 7 )

Type material. Holotype: Male. AUSTRALIA: Queensland: Black Braes National Park: Pine Yards, Einasleigh R. 8.31km E of homestead, 860m, sandy creek bed near permanent waterhole, malaise. 10-13. XI. 2001, D. Yeates, C. Lambkin, N. Starick & J. Hamilton. GPS: 19°32’13”S 144°17’01”E ( ANIC _29:005921) ( ANIC). Condition: Ex - alcohol, critical point dried, micro pinned left side to pith block, reasonable condition.

Paratypes: 6 Males, 13 Females. AUSTRALIA: Queensland: 23 1Ƥ same data as holotype; ( ANIC _29:005922, 29:005923) ( ANIC): Queensland, 33 8Ƥ Queensland, Gorge Creek Station: 27.5 km W of Black Braes Homestead. 951m, dry, stony creek bed, Malaise; D. Yeates C. Lambkin N. Starick & J. Hamilton; GPS: 19°32’53”S 143°56’33”E ( ANIC). ( ANIC _29:005924, 29:005926, 29:005927, 29:005930-29:005936, 29:006293) ( ANIC): 13 3Ƥ Queensland, Black Braes National Park, Wolfram Springs, 13.7km WSW of homestead, 980m, in ravine, over waterholes, Malaise. D. Yeates C. Lambkin N. Starick & J. Hamilton; GPS 19°34’58”S 144°04’57”E ( ANIC _29:004656, 29: 0 0 4657, 29:005928, 29:005929) ( ANIC):1Ƥ Queensland, Black Braes National Park, Log Springs, 17.5km WNW of homestead, 980m, over standing water, Malaise, D. Yeates C. Lambkin N. Starick & J. Hamilton; GPS: 19°34’01”S 144°02’24”E ( ANIC _29:005925) ( ANIC). Addition label on all specimens: Anabarhynchus Macquart, 1848 ; det. J. Hamilton 10. iv. 2002.

Diagnosis. Frons raised; frontal setae short and semi-erect; mid-frons with dark-brown mark angled towards upper frons near meeting along mid-line; male frons width at anterior ocellus 4.6-5.4x ocellus, female 3.2-3.8x ocellus. Grey scutum with indistinct lines. Wing cell m3 open. Postspiracular pile present; katepisternum and prosternal furrow without pile. Forefemur 2 pd, 1 pv, 1 av; mid-femur,1 av, hind femur 1 av; male femora yellowbrown with basal fore and hind femora mottled dark-brown, female femora blackish-brown. All femora with sparse appressed pale pile admix with sparse short black setae predominantly on apical half. Abdominal integument in male’s yellow-brown, in female’s dark-brown.

Description. Male: Body length: 8mm. Wing length: 6mm. Head. Integument black. Frons raised; frons width at anterior ocellus 2.6x ocellus; ocellar tubercle flat; antennal base positioned low on frons; face and lower frons slightly protruding; parafacials grey, lower frons grey, mid-frons with pair dark-brown marks adjacent to eye edge to angled towards upper frons, meeting at mid-line, darken and lighten depending on angle of view; upper frons grey with two rows of semi-erect short black setae; over brown markings of mid-frons setae angle towards eye margin, setae length at lower frons near half length of scape. Scape length 2.3x width; scape and pedicel grey, 1st segment of flagellum brown with a few short black setae to basal dorsal third, flagellar style black one third that of flagellum. Occiput convex with grey pubescence margin of eye yellowish-grey with several indistinct rows of black macrosetae, 28 each side; postocciput area to gena grey, densely supplied with long, pale, hair-like pile. Palpus pale brown, with pale pile; labellum grey, prementum setae black. Thorax. Integument black. Scutal chaetotaxy black (pairs): np, 4; sa, 2; pa, 1; dc, 2; sc, 2. Scutum with thin brown dorsal line within broader grey band, margined by narrow lighter grey line, irregular brownish grey marks perimeter laterally grey raised areas of pre and post-transverse suture; scutal surface with sparsely short, appressed, pale setae. Postspiracular pile present; katepisternum and prosternal furrow without pile; pleuron and coxae with grey pubescence; coxae with long pale setae admixed black macrosetae. Wing. Cell m3 open; hyaline with brown tint, veins and stigma brown, apical half infuscate along veins; costal setae beyond humeral cross-vein biserially arranged. Haltere. Pedicel yellow-brown; scabellum white. Legs. Forefemur 2 pd distributed over apical half, 1 pv, weak, 1 av weak; mid-femur, 1 av weak, hind femur 1 av macrosetae; forefemur basally brownish grey, apical half mottled half yellow; mid-femur yellow, hind femur similar to forefemur though yellow-brown area more extensive; all femora with sparse appressed pale pile, admix with sparse short black setae predominantly to apical half; tibiae yellow-brown apically dark, basitarsus dark. Abdomen. Integument mottled yellow-brown and brown. Laterally compressed; tergites dorsal surfaces yellow-brown without pubescence; tergites 2-4 posterolaterally grey with erect pale pile; tergites and sternites 2-5 posterior marginal bands bright white when viewed posteriorly, grey anteriorly; all sternites similar to tergites; epandrium yellow-brown covered with yellow-grey pubescence, marginal edges with short pale pile; gonocoxites yellow-brown. Terminalia. Paratype male (ANIC_29:005936): epandrium ( Fig. 7 a), three times wider than long, narrowing posteriorly, setae on posterior margin. Gonocoxite ( Fig. 7 b), semi-spherical slightly longer than wide when viewed ventrally; posterior ventral edge with a broad low lobe like outer gonocoxal process, posterior margin with long black setae; gonocoxites connected by hypandrium and thin membrane along ventromedial plane; gonocoxal apodeme long and sclerotised. Inner gonocoxal process ventrally and inwardly directed apical ventral edge with several strong ventral and inwardly directed setae. Gonostylus elongate with setae on dorsal and ventral edges. Ventral lobe absent. Aedeagus ( Fig. 7 c, d): distiphallus short, curved ventrally. Parameral sheath darkly sclerotised. Dorsal apodeme of parameral sheath broadly triangular with sclerotised lobe to lateral edge. Ventral apodeme narrow, flaring apically. Lateral ejaculatory apodeme dorsally broad narrowing ventrally, band-like. Ejaculatory apodeme long extends well beyond dorsal apodeme, cylindrical, apically ends with lateral flanges.

Variation. Male. Body length range: 7–8.5mm. Wing length range: 6–8mm. Head: Frons width at anterior ocellus 4.6–5.4x ocellus; forefemur, 1–4 pd distributed over apical two third, 1–4 pv, weak 1 av weak; mid-femur, 1– 2 pd over apical half, 1–3 pv, often weak 1–2 av often weak; hind femur, 1–2 av macrosetae. Female. Body length range: 7–7.5mm. Wing length range: 6–8.5mm. Frons width at anterior ocellus, 3.2–3.8x ocellus. Forefemur blackish-brown to brown; mid-femur basally half dark brown, apically yellow-brown mottled towards base to all dark-brown; hind femur dark except ventral edge of apical mottled yellow-brown; all femora show variability in mottled colouration. Abdomen integument dark-brown; tergites 2–7 broad anterior blackish-brown bands, with bright light-grey pubescence to posterolateral of tergite 2–4 tergite, tergites 5–7 dull grey, with erect black setae; sternite 2 has bright white-grey pubescence. Paratype female (ANIC_29:005925): sternite 8 ( Fig. 7 f), rounded square shape, darkly sclerotised; anterior edge with slight concavity with thinly sclerotized lobed shaped area centre to this concavity; posterior with squarish bi-lobe inserted into main sclerite, elongate slightly depressed area along mid-line. Furca ( Fig. 7 e), frame narrowing posteriorly with a pair of small internal struts at middle; anterior beam broad with lateral directed projections; anteroventral lobe connected to beam at two mid-lateral points, middle anterior edge strongly indented.

Etymology. The specific epithet is derived to honour the original Aboriginal inhabitants of the land where the type specimens were collected whose tribal name is ‘Ewamin’ ( Tindale, 1974).

Comments. From a series of specimens collected from the Black Braes National Park, inland from Townsville, Queensland, in November.

Keys to couplet 82 in Lyneborg (2001); readily separated from A. gascoyne Lyneborg by having a grey scutum with indistinct markings. Fore femur with 2 pv macrosetae and midfemur without pd and pv macrosetae, but with 1– 2 av weak macrosetae.

Female sternite 8 posterior lobe broadly confluent with the main sclerite with an elongate slightly depressed area midline, subsequently A. ewamin sp. n., is placed in the kroeberi species-group.

kroeberi species-group. Forefemur Mid-femur H/femur

pd pv av pd pv av av

A. boharti Lyneborg 2–3 1 1 1–2 1 1 1–2

A. collessi Lyneborg 2 0 0 0 0 0 1

A. ewamin sp. n. 2 1 1 0 0 1 1

A. gascoyne Lyneborg 2 0 1weak 1 1 1 1–2

A. kroeberi Lyneborg 2 0 0 1 1 0 2

A. pilbara sp. n. 2 0 0 1 0 0 1–2

A. ravenshoensis sp. n. 1–2 2 1 1 1 1 1

A. tribulationensis sp. n. 2 1–2 0 1 4–7 1–2 1–2

A. weipaensis sp. n. 3–4 0 0–2 1 0 0 1–2

Figures

FIGURE 5. Anabarhynchus ewamin sp. n., holotype male, (ANIC _ 29: 005921): lateral view.

FIGURE 6. Anabarhynchus ewamin sp. n., holotype male, (ANIC _ 29: 005921): oblique view.

FIGURE 7. Anabarhynchus ewamin sp. n., paratype male terminalia (ANIC _ 29: 005936): A, epandrium, left dorsal view, right ventral view; B, gonocoxite left dorsal view, right ventral view; C, aedeagus, dorsal view; D, aedeagus, lateral view; paratype female (ANIC _ 29: 005925): E, furca; F, female sternite 8, ventral surface. Scale line = 0.5 mm.